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Plant Breeding: Quiz 2
University of Guelph MBG*4160
209
Microbiology
Undergraduate 4
11/06/2015

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Term
Additive dominance
Definition
A form of incomplete dominance. The heterozygoute is at a midpoint between the homozygotes. Example: the Waxy gene in maize.
Term
Additive effect (a)
Definition
When two genes produce the same effect, but effects are additive if both are present. The difference between two homozygotes.
Term
Additive variance
Definition
A component of genetic variance. Variation passed intect from parent to progeny. Variation in breeding value.
Term
Adventitious embryony
Definition
A type of apomixis. The embryo develops from cells in tissues outside the embryo sac: ovule, nucellus, or integument cells. Most require pseudogamy for seed set. Fertilization occurs, but the nucellar embryo has a faster growth rate than the zygotic embryo. Polyembryonic seeds are produced. The endosperm is generated by fertilization.
Term
Allele
Definition
Different versions of a gene at the same locus.
Term
Allele frequency
Definition
The frequency of a certain allele in a population.
Term
Allopolyploid
Definition
A type of polyploid.
Term
Anther
Definition
The male reproductive organ of flowers. Contains the microsporangia. Each species has a different number of anthers, but the number is constant to each species.
Term
Antipodal nuclei
Definition
Cells in the embryo sac. Three are at the chalazal end of the embryo sac.
Term
Apomixis
Definition
A form of asexual reproduction. Production of seeds from somatic cells of the maternal parent. For some plants, psuedogamy is necessary to induce apomixis. Progeny is genetically identical to the mother. Example: citrus, bluegrass. Four types of apomixis: adventitious embryony, apospory, mitotic diplospory, and meiotic diplospory.
Term
Apospory
Definition
A type of apomixis. An embryo sac develops from cells that differentiated from the nucellus after formation of the megasporocyte. The embryo sac is diploid, so it grows faster than the haploid embryo sacs (not delayed by meiosis). The endosperm is formed by pollination from pseudogamy.
Term
Approach method
Definition
A method of artificial hybridization in wheat. The male spike is placed inside the glycine bag, kept in a drinking straw filled with water.
Term
Artificial hybridization in Brassicas
Definition
Wind is a more important pollinator than insects. Flowers are indeterminate, and the stigma is receptive 3 days before and 3 days after the flower opens. Pollen can be stored for 4 - 5 weeks.
Term
Artificial hybridization in carrots
Definition
The flowers are protandrous: pollen is shed before stigmas are receptive. The plants are fastened below the flowers and placed in a cage with flies that pollinate the flowers.
Term
Artificial hybridization in soybeans
Definition
Flowers are typically self-pollinated. Natural cross-pollination is about 1% from insect vectors. The flower has 5 petals: the standard, two wing petals, and two keel petals.
Term
Artificial hybridization in wheat
Definition
Two methods: Go-Go method and Approach method.
Term
Asexual reproduction
Definition

Vegetative reproducction

Reproduction through plant parts other than the seed. Includes: bulb, corm, tuber, rhizome, stolon, micropropagation, and somatic embryogenesis. Apomixis, parthenogenesis, and artificial means such as cutting, grafting, layering, and tissue culture are also forms of asexual reproduction.

Term
Autopolyploid
Definition
A type of polyploid.
Term
Awned
Definition
There are long spikes, awns, on the lema of wheat or barley.
Term
Axial cell
Definition

Apical cell

One of the cells formed in the first division of the zygote. It divides to form the embryo.

Term
Basal cell
Definition
One of the cells formed in the first division of the zygote. In dicots it divides to form the suspensor. In monocots it does not divide, and forms the terminal cell of the suspensor.
Term
Basmati rice
Definition
Rice with a good flavour, controlled by a genet. But no-one created the gene! Can you patent it?
Term
Binucleate stage
Definition
The first stage in microgametogenesis. The microspore has cytoplasmic division, producing a generative cell and a tube cell. Most pollen is released in this stage.
Term
Both self- and cross-pollination
Definition
Some species have both self- and cross-pollination, with about 50% of each. Example: cotton, pigeon pea.
Term
Brassica campestris
Definition
Highly self-incompatible. Self-pollinated seed is produced with difficulty.
Term
Brassica hirta
Definition
Highly self-incompatible. Self-pollinated seed is produced with difficulty.
Term
Brassica jucea
Definition
Self-fertile. 80% of seed is from self-pollination. This number is smaller when insect pollinators are abundant.
Term
Brassica napus
Definition
Self-fertile. 80% of seed is from self-pollination. This number is smaller when insect pollinators are abundant.
Term
Breeding Field Crops
Definition
A book by John Pullman.
Term
Breeding Value
Definition
The value of an individual as judged by the mean value of the progeny.
Term
Broad sense heritability
Definition

Genotypic variation / Phenotypic variation

An estimate of heritability. Measures the proportion of total variance that is genetic. Found using selfed progeny and progeny-parent regression heritability estimates.

Term
Bulb
Definition
A form of asexual reproduction. A large bud with a small stem at the lower end; buds in the axils of fleshy, scale-like leaves permit the bulb to be divided for asexual reproduction. Example: onion, garlic, tulips, lilies.
Term
Cellular endosperm development
Definition
The endosperm divides normally with cytokinesis; there is no free nuclear stage.
Term
Centre of origin
Definition
The origin of a cultivated species. Has wild species in addition to landraces. High level of genetic diversity.
Term
Chalazal end
Definition
The end of the embryo sac opposite from the micropylar end. The antipodal cells are on this end.
Term
Chasmogamy
Definition
A device that promotes out-crossing. Flowers are open at anthesis.
Term
Chi-square (Χ2)
Definition
A test of goodness of fit. The smaller the value, the more likely the hypothesis is correct.
Term
Chiasma
Definition

Plural: chiasmata

The point where two homologous, non-sister chromatids exchange genetic material during crossing over.

Term
Chloroplasts
Definition
One of the places in the plant cell where DNA is kept. Chloroplast DNA is inherited maternally in most angiosperms, from both parents in some angiosperms, and paternally in a few angiosperms and most gymnosperms.
Term
Chromosome
Definition
A condensed strand of DNA.
Term
Cleistogamy
Definition
A device that promotes self-pollination. Flowers are closed at anthesis. Example: soybean, barley, wheat, oats.
Term
Colchicine
Definition
A chemical that doubles the number of chromosomes.
Term
Complementary action
Definition
When two non-allelic genes are required to produce a single effect.
Term
Complementation
Definition

Duplicate recessive

A type of epistasis

aa is epistatic to B and b

bb is epistatic to A and a

Produces 9:7 ratios

Term
Complete dominance
Definition
When the heterozygote shows the same phenotype as one of the homozygotes.
Term
Complete flower
Definition
Has all four floral organs: sepals, petals, stamens, and pistils.
Term
Corm
Definition
A  form of asexual reproduction. Resembles a bulb in size and form, but has different internal structure. Leaves are thin and small. Roots develop from the lower surface of the stem. Example: crocus, gladiolus.
Term
Cross-pollinated species
Definition

Allogamy

Have devices to promote outcrossing: chasmogamy, dioecious flowers, monoecious flowers, pin and thrum flowers, protandry, protogyny, protective film over the stigma, self-incompatibility, and male sterility. Often perennial. May also have asexual modes of reproduction. Individuals are heterozygous at many loci. Reduced performance when self-pollinated; inbreeding depression due to deleterious alleles in the population. Wild populations are heterogenous. Cultivated varieties are either heterogeneous or homogenous.

Natural self-pollination rates: 0% - 5%

Examples: corn, alfalfa, carrot, onion, rye, sunflower

Term
Crossing over
Definition
A source of diversity from meiosis. Formation of a chasma and exchange of genetic material between chromosomes. Results in recombinant chromosomes. Genotype frequencies are unusual.
Term
Cytoplasmic DNA
Definition
DNA in the mitochondria or chloroplast. Circular DNA, distinct from nuclear DNA. Usually inherited uniparentally.
Term
Cytoplasmic male sterility
Definition
Male sterility controlled by the cytplasm, with many influences from nuclear genes, inhibiting normal development of anthers.
Term
Davis
Definition
A soybean variety resistant to Frogeye leaf spot.
Term
Dioecious
Definition

A device that promotes out-crossing in imperfect flowers. Male and female reproductive parts are on separate plants.

Example: date plum, papaya, asparagus, hemp, hops

Term
Diploid
Definition
Two chromosome sets, one from each parent. Chromosomes are in homologous pairs. May be homozygous or heterzygous.
Term
DNA
Definition
A long, double-helix molecule with 2 complementary strands with opposite polarity.
Term
Dominance
Definition
The dominance of alleles determines the phenotype of an organism.
Term
Dominance variance
Definition
Genetic variance that results from allelic interactions within loci.
Term
Dominant epistasis
Definition

A type of epistasis.

A is epsitatic to B and b

Produces a 12:3:1 ratio

Term
Double fertilization
Definition
Unique to angiosperms. The egg cell is fertilized by a sperm cell, resulting in a diploid zygote. The polar nuclei are ferilized by the second sperm, resulting in the endosperm.
Term
Duplicate action
Definition
When either of two genes produce a similar effect.
Term
Duplication
Definition

A type of epistasis

A is epistatic to B and b

B is epistatic to A and b

Prouces a 15:1 ratio

Term
Durum wheat
Definition

Triticum turgidum

2n = 4x = 28 AABB

One of the parents of triticale.

Term
Egg
Definition
A cell in the embryo sac. At the micropylar end. It is fertilized by a sperm cell to produce the zygote.
Term
Embryo
Definition
A diploid component of a seed. Develops from the axial cell of the zygote. In dicots it has four stages of development: globular, heart, late heart, and torpedo stage.
Term
Embryo sac
Definition

Megagametophyte

Formed from the megapore in megagametogenesis. IT consists of seven cells and eight haploid nuclei: one egg, two polar nuclei (in one cell), two synergids, and three antipodal nuclei. Some species have more or less than the usual eight nuclei. It is surrounded by integuments, with the micropyle at one end. The micropylar end is near the micropyle, and the chalazal end is opposite from teh micropyle.

Term
Endosperm
Definition
Forms from fusion of a sperm and the polar nuclei. Usually triploid, but can be 2n through to 15n. It draws nutrients from adjacent tissues. It either has nuclear development or cellular development. In monocots and endospermic dicots (such as castor beans0, the endosperm is generally no used until germination). In non-endospermic dicots, the endosperm reserve is depleted and reorganized into bulky, nutrient-rich cotyledons during seed development.
Term
Epistasis
Definition
Expression of a gene at one locus affects the phenotypic expression at another locus. Incldues recessive epistasis, dominant epistasis, suppression, complementation, and duplication.
Term
Epistatic variance
Definition
Genetic variation created by interactions between loic. Interactions are not passed intact from parent to progeny. Difficult to select for.
Term
Environmental variance
Definition
Variance due to the environment. Due to the inability to treat all genes equally.
Term
Essentials of Plant Breeding
Definition
A book by Rex Bernardo.
Term
Frogeye Leaf Spot (FLS)
Definition
A disease in soybeans caused by a fungus in hot humid soybean regions such as Brazil, China, Nigeria, and southern US. Causes yield losses 10 - 50%. Resistance is controlled by one or two genes. Susceptible plants have large, spreading, fertile lesions. Resistatn plants have no lesions, and small sterile flecks.
Term
Full-sib progeny
Definition

A type of progeny-parent regression

b = narrow-sense heritability

Term
Gametophytic self-incompatibility
Definition
Self-incompatibility where the genotype of the haploid gametes determines the compatibility reaction. If the allele in the pollen matches one in the female plant, then fertilization fails.
Term
Gene
Definition
A stretch of DNA that transcribes a functional RNA or enables transcription of a protein.
Term
Gene bank
Definition
A source of genetic diversity. Accessions are categorized by geographical source and phenotypic traits. Canada's main gene bank used to be in Ottawa, but was moved to Saskatoon in 1988.
Term
Generative cell
Definition
One of the cells formed in the binucleate stage of microgametogenesis. It divides to form two sperm cells in the trinucleate stage.
Term
Genetic advance
Definition
The change in mean of a population from one generation cycle of selection.
Term
Genetic male sterility
Definition
Male sterility manifested through action of nuclear genes, inhibiting normal development of anthers.
Term
Genetic variance
Definition
The variation due to genetic differences among plants. Consists of additive, dominance, and epistatic variance.
Term
Genetics
Definition
The study of genes through their variation, or the study of inheritance.
Term
Genome
Definition
The entire genetic information of an organism.
Term
Genotype by environment (G x E) variance
Definition
Variance due to genotypes responding differently to different environments. Reduces the association between genotypic and phenotype values. May cause selections from one environment to perform poorly in another environment. Minimized by using many environments and many replications.
Term
Genotype frequency
Definition
The frequency of a certain genotype in a population.
Term
Germplasm Resource Information Network (GRIN)
Definition
A source of information on accessions in gene banks. You can order accessions for free from the Canada gene bank.
Term
Globular stage
Definition
The first stage of embryo development. The embryo is sphere shaped.
Term
Go-Go method
Definition
A method of artificial hybridization in wheat. The male spike is shaken inside the opening bag containing the female spike. The bag is then resealed.
Term
Half-sib progeny
Definition

A type of progeny-parent regression.

2b = narrow-sense heritability

Term
Hardy-Weinberg Equilibrium
Definition
Described by G.H. Hardy and W. Weinberg in 1908. Frequencies of alleles and genotypes stay constant if the population is large, randomly mating, absent of mutations, absent of selection, absent of migration.
Term
Heart stage
Definition
The second stage of embryo development. Three major regions of mitotic activity: two on either side of the top of the embryo, and one near the suspensor. These develop into the cotyledons and radicle.
Term
Heritability (h2)
Definition
The proportion of the phenotypic variation that is due to genotype. Estimates of heritability can be used to predict selection gain. Includes broad sense and narrow sense estimates.
Term
Heterozygous
Definition
Having different alleles on two homologous chromosomes.
Term
Hooded
Definition
The lema of barley or wheat has a "hood" over it.
Term
Homologous chromosomes
Definition
Chromosomes in a diploid organism which carry the same gene loci. May be homozygous or heterozygous.
Term
Homozygous
Definition
Having the same allele on both homologous chromosomes.
Term
High leaf starch (hls)
Definition
A mutation in alfalfa causing increased activity of invertase, leading to starch build-up in the leaf. It is transmitted paternally more than it is transmitted maternally; this established that in Medicago species plastids are inherited paternally. It is studied using Restriction Fragment Length Polymorphism.
Term
Hybrid vigour
Definition
When the heterozygote is better than either homozygote.
Term
Incomplete dominance
Definition
Includes additive, partial, and over-dominance.
Term
Incomplete flower
Definition
Lacks one or more floral organs: sepals, petals, stamens, or pistils.
Term
Independent assortment of chromosomes
Definition
A source of diversity from meiosis. Different chromosomes are independently separated from one another during gamete formation.
Term
Inhibiting action
Definition
When one gene inhibits the effect of another gene at another locus.
Term
Integuments
Definition
Surround the embryo sac. Has a small opening at one end, the micropyle. After fertilization, develops into the testa.
Term
Invertase
Definition
A protein that exports sucrose from the cell. In high leaf starch alfalfa mutants it has increased activity, resulting in starch build-up in the leaf.
Term
Keel petal
Definition
A petal in soybean flowers. There are two keel petals.
Term
Landrace
Definition
A cultivated form that evolved from a wild population, which is endemic to an area with origins going back 100s of years, well adapted to the environment, a mixture of types.
Term
Late heart stage
Definition
The third stage of embryo development. Elongation of regions of mitotic activity, defining cotyledons.
Term
Law of Independent Assortment
Definition
Alleles at a locus segregate independently from alleles at another locus, when gametes are formed.
Term
Law Segregation
Definition
Alleles of a gene separate randomly when gametes are formed.
Term
Lee
Definition
A soybean vareity susceptible to most races of Frogeye leaf spot.
Term
Linkage
Definition
Genes are on the same chromosome. Produces unusual segregation ratios.
Term
Locus
Definition
The location of a gene on a chromosome.
Term
Male sterility
Definition
A device that promotes out-crossing. Includes genetic and cytoplasmic male sterility.
Term
Masking action
Definition
When one gene hides the effect of another gene at another locus, when both are present.
Term
Mean (μ)
Definition
A measure of central tendency, used when studying quantitative traits in populations.
Term
Megagametogenesis
Definition
The megaspore undergoes division to form seven cells and eight nuclei that form the embryo sac.
Term
Megaspore
Definition
A haploid cell formed via meiosis of the megasporocyte. Arranged in a tetrad cluster or linear arrangement. Four are produced per megasporocyte, but three degenerate.
Term
Megasporocyte
Definition

Megaspore mother cell

Undergoes meiosis to form four megasporogenesis to form four megaspores.

Term
Megasporogenesis
Definition
The megasporocyte undergoes meiosis to form four megaspores within the nucellus.
Term
Meiosis
Definition
Diploid cells divide to produce haploid cells. It produces diversity from independent assortment of chromosomes, and from crossing-over. Halves the number of chromosomes during production of sex cells. There is independent segregation of genes.
Term
Meiotic diplospory
Definition
A type of apomixis. The megasporocyte differentiates from the nucellus, but has a failure in meiosis I, resulting in a diploid megaspore. Development of an embryo from parthenogenesis; pollination is not required.
Term
Microgametogenesis
Definition
Development of pollen grains from microspores. Includes the binucleate and trinucleate stages.
Term
Micropropagation
Definition
A form of asexual reproduction.
Term
Micropylar end
Definition
The end of the embryo sac near the micropyle. The egg and synergids are on this side.
Term
Micropyle
Definition
A small hole in the integuments on the micropylar end of the embryo sac. The pollen tube enters through this opening.
Term
Microsporangia
Definition

Pollen sacs

Microsporogenesis occurs here.

Term
Microspore
Definition
A haploid cell formed via meiosis of the microsporocyte in microsporogenesis.
Term
Microsporocyte
Definition

Microspore mother cell

Undergoes meiosis in microsporogenesis, forming 4 microspores.

Term
Microsporogenesis
Definition
The microsporocyte undergoes mieosis to form 4 microspores within the microsporangia of the anther.
Term
Mitochondria
Definition
One of the places in the plant cell where DNA is kept. Mitochondrial DNA is inherited maternally.
Term
Mitosis
Definition
Diploid cells divide to produce diploid cells. Chromosomes are partitioned into daughter cells during somatic division.
Term
Mitotic diplospory
Definition
A type of apomixis. The megasporocyte is inhibited from entering meiosis. The embryo sac is diploid, but is identical in structure to the haploid embryo sac. The embryo is formed from parthenogenesis; pollination is not necessary. The endosperm is also produced by parthenogenesis, but in some cases pseudogamy is required form the endosperm.
Term
Modifying action
Definition
When one gene inhibits the effect of another gene at another locus.
Term
Monoecious
Definition
A device that promotes out-crossing in imperfect flowers. Male and female reproductive parts are on separate flowers on the same plant. Example: maize
Term
Mutation
Definition
Genes or chromosomes changing from one allele to another.
Term
Narrow sense heritability
Definition
An estimate of heritability. The proportion of variance that is additive. The proportion which can be selected for. Found using full-sib and half-sib progeny-parent regression method.
Term
Nucellus
Definition
Megasporogenesis occurs here. Devlops into the perisperm in some species.
Term
Nuclear endosperm development
Definition
The endosperm undergoes several free nuclear divisions prior to cell wall formation.
Term
Nucleus
Definition
One of the places in the plant cell where DNA is kept. Nuclear DNA is inherited maternally and paternally.
Term
Over-dominance
Definition
A type of incomplete dominance. The homozygote is greater than either heterozygote.
Term
Parthenogenesis
Definition
A form of asexual reproduction. An embryo develops without fertilization, resulting in haploid progeny. No pollination is necessary.
Term
Partial dominance
Definition
A type of incomplete dominance. The homozygote is between both homozygoutes, but is not at the midpoint.
Term
Peking
Definition
A soybean variety that is resistant to Frogeye leaf spot.
Term
Perisperm
Definition
Develops from the nucellus in some species. Assumes the role of the endosperm. Diploid, derived from the mother plant. Example: coffee.
Term
Phenotype
Definition
Determined by the dominance of alleles in an organism.
Term
Phenotypic variance
Definition
Genetic variance + Environmental variance + G x E interaction
Term
PI 51461
Definition
A soybean variety resistant to Frogeye leaf spot.
Term
Pin and thrum
Definition
A device that promotes out-crossing in imperfect flowers. Example: primula.
Term
Plant DNA
Definition
Found in the nucleus, mitochondria, and chloroplasts.
Term
Pleiotropic genes
Definition
When a single gene may have more than one effect.
Term
Polar nuclei
Definition
A component of the embryo sac. Two nuclei in one cell. Near the centre of the embryo sac. It is fertilzied by a sperm cell to form the endosperm.
Term
Pollen grain
Definition

Microgametophyte

Formed from microspores in microgametogenesis.

Term
Pollination
Definition
Pollen grains are transferred into the stigma by a variety of vectors: insects, animals, wind. Many species enter trinucleate stage at this point, producing two sperm cells. The pollen grain absorbs water and nutrients from the stigma surface, and the pollen tube emerges and grows through the stigma and style to the embryo sac, and enters the ovule through the micropyle. It penetrates a synergid, and deposits sperm cells.
Term
Polyembronic seeds
Definition
Seeds with more than one embryo. Created in adventitious embryony; there are a number of nucellar embryos in different stages, with or without a zygotic embryo.
Term
Polyploid
Definition
Originate as autopolyploids or allopolyploids.
Term
Population
Definition
A group of interbreeding individuals that exist together in time and space. Distribution is measure with mean and variance.
Term
Pre-breeding
Definition
Creates genetics that you cannot get directly. There are seveal processes before getting the final product. Breeding with landraces or wild species transfers unwanted traits, which are removed in pre-breeding.
Term
Princess
Definition
A variety of dry bean Johannsen used in his experiments. Continuous breeding had no result because it was a pure-line.
Term
Progeny-parent regression
Definition
Calculating the regression coefficient of a line of parental vs. progeny phenotype. A higher number means a higher heritability. There are three types: selfed progeny, full-sib progeny, and half-sib progeny.
Term
Progeny testing
Definition
Learning the genotype of a plant by growing and observing characteristics of its progeny.
Term
Protandry
Definition
A device that promotes out-crossing. Stamens develop before pistils.
Term
Protective film over the stigma
Definition
A device that promotes out-crossing in perfect flowers.
Term
Protein
Definition
The product of genes. Transcribed based on the sequence of RNA strands.
Term
Protogyny
Definition
A device that promotes out-crossing. Stamens develop after pistils.
Term
Pseudogamy
Definition
When pollination and pollen tube development is necessary to induce apomixis.
Term
Pure line
Definition
The progeny of a single plant, self-fertilized over many generations. All plants are homozygous and homogenous.
Term
Pure Line Theory
Definition
Postulated by Wilhelm Johannsen in 1903. Continuous inbreeding leads to homozygosity. Variation within pure lines is caused by the environment only. Selection within a pure line is ineffective; all plants are genetically identical.
Term
Qualitative trait
Definition
Controlled by a few loci. Expression is not influenced by the environment. Phenotypes can be divided into discrete classes. Studied by analyzing the phenotypic ratios and inheritance patterns. Can show complete or incomplete dominance, and epistasis. Example: awned vs. hooded wheat and barley.
Term
Quantitative trait
Definition
Controlled by many loci. Expression is influenced by the environment. Phenotypes overlap and/or display continuous variation. Genotype cannot be determined from phenotype. Studied using measures of central tendency in populations. When environmental variation is eliminated, it can appear qualitative; Mendelian segregation patterns are followed. Example: yield in wheat.
Term
Recessive epistasis
Definition

A type of epistasis

aa is epistatic to B

Produces a 9:3:4 ratio

Term
Reciprocal cross effect
Definition
Sometimes a different genotype is produced from different combinations of the sexes of the parents. Example: A female x B male vs. B female x A male, produce different phenotypes in the offspring. Indicates that genes are on mitochondrial or chloroplast DNA. Studied with Restriction Fragment Length Polymorphism. Example: high leaf starch mutation.
Term
Recombinant
Definition
Gene trait combinations that are a result of crossing-over.
Term
Regression coefficient (b)
Definition
The slope of the line made in a progeny-parent regression method for estimating heritability.
Term
Response to selection (R)
Definition

ckh2Ve

k is standardized selection differential

c is the parental control factor

To improve R, you can increase selection intensity, variability, or heritability (reduce errors in measurement of phenotype).

Term
Restriction Fragment Length Polymorphism (RFLP)
Definition
A marker used to follow transmission of plastids. Used to study high leaf starch mutation in alfalfa.
Term
Rhizome
Definition
A form of asexual reproduction. An underground horizontal stem that shoots and roots at nodes. It can be mistaken for a root when it grows underground. Example: bamboo, sugarcane, banana.
Term
RNA
Definition
A single-stranded molecule. Translated from genes. Can enable transcription of a protein.
Term
Rye
Definition

2n = 2x = 14 RR

One of the parents of triticale.

Term
Seedless watermelon
Definition
Triploid watermelon are sterile and produce no seeds. A diloid watermelon is treated with colchicine to produce a tetraploid watermelon, which is then backcrossed with diploid watermelon to create triploid progeny.
Term
Selection differential (S)
Definition
The mean phenotypic value of all the individuals selected as a derivation from the population mean.
Term
Selection intensity
Definition
The proportion of the population which the breeder discards.
Term
Self-incompatibility (SI)
Definition
A device that promotes out-crossing. The inability of a plant to produce a zygote by self-pollination. Several mechanisms: pollen fails to germinate on the stigma, pollen tube is inhibited, pollen tube is insufficient length or strength, or the male gamete fails to unite with the egg cell. Can be gemetophytic or sporophytic.
Term
Self-pollinating species
Definition

Autogamy

Have devices to promote self-pollination: cleistogamy, staminal sheaths. Often annual species. Individuals are homozygous at many loci; there is no inbreeding depression. Wild populations are heterogeneous. Cultivated varieties are homogeneous - all individuals have the same genotype. Inheritance follows the Pure Line Theory.

Natural cross-pollination rates: 0% - 5%

Examples: barley, common bean, chickpea

Term
Selfed progeny
Definition

A type of progeny-parent regression

b = broad-sense heritability

Term
Sexual reproduction
Definition
Fusion of male and female gametes, regardless of whether they originate from one or more flowers or plants. Includes cross- and self-pollinating species. Each group has devices to promote their mode of reproduction.
Term
Sperm cell
Definition
Two cells that arise from the generative cell of a pollen grain in trinucleate stage. One fertilizes the egg cell, forming the zygote. The other fertilizes the polar nuclei, forming the endosperm.
Term
Sporophytic self-incompatibility
Definition
Self-incompatibility where the genotype of the male parent determines the compatibility reaction. If either allele in the male parent matches one in the female parent, fertilzation fails, regardless of the genotype of the pollen grain. The genotype of the male parent is marked on the exine.
Term
Somatic embryogenesis
Definition
A form of asexual reproduction. An embryo forms, and balanced hormones are used to produce a plant. Fertilization does not occur.
Term
Staminal sheath
Definition
A device that promotes self-pollination. The stamens are joined, surrounding the pistil. The pistil grows through the pollen, causing self-pollination. Example: cotton.
Term
Standard petal
Definition
A petal of a soybean flower. There is one standard petal.
Term
Stigma
Definition
The floral organ which receives pollen.
Term
Stolon
Definition

Runners

A form of asexual reproduction. An above-ground horizontal stem. Example: strawberry, creeping bentgrass, Bermuda grass.

Term
Suppression
Definition

A type of epistasis.

A is epistatic to B and b

bb is espistatic to A and a

Produces a 13:3:0 ratio

Term
Suspensor
Definition
In dicots it is formed from the basal cell. In monocots it is formed from the axial cell, but its terminal cell is formed from the basal cell. Synthesizes growth factors and transports nutrients to the embryo. Can be unicellular, multicellular, large, small, filamentous, columnar, spherical, or irregular in shape.
Term
Synergids
Definition
Cells in the embryo sac. There are two at the micropylar end. The pollen tube penetrates a synergid and disintigrates after fertilization.
Term
Terminal cell
Definition
Formed from the basal cell in monocots. Becomes part of the suspensor, which is derived from the axial cell.
Term
Test cross
Definition
Crossing a plant with a homozygous recessive individual to identify its genotype.
Term
Testa
Definition

Seed coat

Develop from integuments. Diploid, derived from the mother plant.

Term
Torpedo stage
Definition
The fourthe stage of embryo development. Further elongation results in a bend in the region of the hypocotyl.
Term
Transcription
Definition
The formation of a protein based on the sequence of an RNA strand.
Term
Transgressive segregation
Definition
When progeny fall outside of the range of the parents.
Term
Trinucleate stage
Definition
The second stage in microgametogenesis. In most species it occurs when the pollen grain lands on the stigma. In others, it occurs before pollen is released. The generative cell undergoes division, forming two sperm cells.
Term
Triticale
Definition
The first man-made crop. A cross between durum wheat (4n = 28 = AABB) and rye (2n = 12 = RR), creating a sterile 3n = 21 = ABR individual. Colchicine was used to double the chromosomes, producing 6n = 42 = AABBRR triticale.
Term
Tube cell
Definition

Vegetative cell

One of the cells formed in the binucleate stage of microgametogenesis. Helps the sperm cells reach the egg.

Term
Tuber
Definition
A form of asexual reproduction. A swollen, modified stem that acts as an underground storage organ. Example: potato, Jerusalem artichoke, yam.
Term
Variance (σ2)
Definition
A measure of distribution. Used in studying quantitative traits in populations.
Term
Waxy
Definition
A gene in maize that shows additive dominance.
Term
Wheat
Definition
Has genes transgressed in from GP2 and GP3. It has 21 pairs of chromosomes.
Term
Wilhelm Johannsen
Definition
Postulated the Pure Line Theory in 1903. Used Princess dry beans in his experiments.
Term
Wing petal
Definition
A petal in soybean flowers. There are two wing petals.
Term
Zygote
Definition
A diploid cell formed from fusion of the egg and a sperm. It develops into the embryo of the seed. The first division in the zygote forms the axial and basal cells.
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