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Phylum Annelida
3/12/14 Test
89
Biology
Undergraduate 3
03/11/2014

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Term
A. Metamerism
1. Metamere/Somite
Definition
A. Segmental arrangement of body parts in an animal
1. each segment of a worm
Term
Annuli
Definition
grooves found externally BETWEEN each somite
Term
Septa
Definition
Internal membranes formed from adjacent peritoneum of segments
-form internal division membranes
Term
Tagmatization
Definition
the specialization of body regions in a metameric animal.
Term
Setae
Definition
tiny chitinous hairs that ANCHOR the annelid during locomotion to prevent backward slipping
- can be used for swimming as well in some
- bristles secreted from invaginations of the distal ends of parapodia.
Term
Tagmatization - Class Polychaeta
Definition
have a well-developed head with specialized sense organs, PARAPODIA and NO clitellum.
Term
Parapodia - Class Polychaeta
Definition
paired appendages
- numerous setae project from the parapodia, giving them their class name.
- used in locomotion and respiration
Term
Prostomium - Class Polychaeta
Definition
a lobe tat projects dorsally and anteriorly to the mouth.
- contains sensory structures such as the eyes, antennae, sensory palps, and tentacle.
Term
Pygidium - Class Polychaeta
Definition
the terminal portion of the worm where the anus is found
- not considered a somite.
Term
Peristomium - Class Polychaeta
Definition
the 1st segment of the worm
- surrounds the mouth
- may bear cirri
Term
Cirri - Class Polychaeta
Definition
sensory tentacles.
Term
Epidermis - Class Polychaeta
Definition
a single layer of columnar epitelial cells that secrete a non-living cuticle
Term
Cuticle - Class Polychaeta
Definition
a protective secretion by the epidermis
- may contain luminescent compounds.
Term
Longitudinal Muscles - Class Polychaeta
Definition
muscles on each sides of the body that allow movement in polychaetes
Term
Digestive tract - Class Polychaeta
Definition
A straight tube suspended in the coelom by the mesentaries and septa.
- consists of a pharynx, probiscus, crop, grinding gizzard & long intestine

*Predatory - probiscus and poison glands
*Herbivores/scavengers - jaws to tear
*Deposit-feeders - organic matter they ingest
*Filter feeders - no probiscus, replced by radioles
Term
Radioles - Class Polychaeta
Definition
feathery arms that contain cilia that trap food particle in mucous that is carried/
Term
Casting - Class Polychaeta
Definition
what organic matter is known as after it's digested, passed through the intestine and is released...
Term
Gas exchange - Class Polychaeta
Definition
By DIFFUSION through the body wall and parapodia.
Term
Closed Circulatory System - Class Polychaeta
Definition
Oxygen carried in respiratory pigments instead of blood.
- may make the blood colorless, green or red
- contractile elements consist of a DORSAL vessel on dorsal side of the digestive tract that pumps blood ANTERIORLY and a VENTRAL vessel that pumps blood POSTERIORLY
Term
Extra Info- Class Polychaeta
Definition
Polychaetes that live in rich organic substances can absorb as much as 40% of their energy requirements through the body wall
- occurs in other animals but usually only to the extent of 1%
Term
Nervous System - Class Polychaeta

(Each segment acts separately from, but is closely coordinated with, neighboring segments...)
Definition
Consists of a pair of suprapharyngeal ganglia ("brain")which connect to a pair of subpharyngeal ganglia by circumpharyngeal connectives that run dorsoventrally along either side of the pharynx.

A double, ventral nerve cord runs from the subpharyngeal ganglia the length of the worm.
Term
Nervous System (cont.) - Class Polychaeta
Definition
Segmental ganglia - coordinate swimming and crawling movements in isolated segments.

Lateral nerves emerge from each segmental ganglion.
- supplies the body wall musculature & other structures of that segment.

Sensory structures in the head - 2-4 eyes on the surface of the prostomium.
- simple to complex w/cornea, lens, & vitreous humor.
Term
Nuchal organs - Class Polychaeta
Definition
a pair of ciliated sensory pits in the head region acting as CHEMORECEPTORS for food detection
Term
Statocysts - Class Polychaeta
Definition
located on head region for equilibrium and balance
Term
Tactile Receptors - Class Polychaeta
Definition
ciliated tubercles, ridges and bands that cover the body wall
Term
Nephridia - Class Polychaeta
Definition
accomplish the excretion of nitrogenous wastes and help maintain the osmotic balance
Term
Protonephridia - Class Polychaeta
Definition
bulbular end projecting through the anterior wall of the adjoining segment and the opposite end opening on the segment to the outside through a nephridiopore
Term
Nephridiopore - Class Polychaeta
Definition
an opening on the nephridia
Term
Metanephridium - Class Polychaeta
Definition
Consists of a nephrostome that projects through an anterior septum into the coelom of an adjacent segment.

At the other end in the segment containing extensively coiled tubules, with a portion dilated into a bladder, it opens to the outside through a nephridiopore.

(Most common type of nephridia in polychaetes)
Term
Nephrostome - Class Polychaeta
Definition
an open, ciliated funnel
Term
Regeneration - Class Polychaeta
Definition
they can replace lost parts, and some species have break points that allow worms to sever themselves when a predator grabs them.
- lost segments are later regenerated...
Term
Reproduction - Class Polychaeta
Definition
MOST ARE DIOECIOUS
-some reproduce asexually through budding or transverse fission.

- fertilization is external
- trochophore larva bud segments anterior to the anus
- Larva settles to the substrate and adds newer segments posteriorly
- ANTERIOR END OF THE WORM IS THE OLDEST.
- MANY POLYCHAETES LACK A LARVAL STAGE AND HAVE DIRECT DEVELOPMENT.
Term
Epitoky - Class Polychaeta
Definition
the formation of a reproductive individual (EPITOKE) that differs from the nonreproductive form of the species (an ATOKE)
Term
Epitoke - Class Polychaeta
Definition
Frequently has a body that is modified into 2 body regions.
- Anterior are normal where the posterior segments are enlarged and filled with with gametes.
- also may have modified parapodia for more efficient swimming.

All gametes are released at a certain time of the year and fertilized eggs develop into trochophore larva and develop into nonreproductive adults...
Term
Class Clitellata - overview
Definition
-includes the earthworms and leeches
-put into same class because of the presence of a clitellum
-monoecious
-presence of few to no setae
-Subclass Oligochaeta - Earth Worms
-Subclass Hirudinea - Leeches
- BOTH LACK PARAODIA
Term
Clitellum
Definition
cocoon-secreting structure
Term
Subclass Oligochaeta - overview
Definition
-MOST ARE TERRESTRIAL but many live in freshwater.
-some are MARINE or are found in estuaries
-have setae, but aren't as numerous as the polychaetes.
-OLIGOCHAETA mean "few long hairs"
-MOST FAMILIAR >>> the Earthworm or "Lumbricus terrestris"
-LACK PARAPODIA!!!!
Term
Earthworm scientific name
Definition
Lumbricus terrestris
Term
Prostomium
Definition
a small lobe in the front of the mouth that lacks sensory appendages.
Term
Clitellum
Definition
Usually somites 32-37, swollen into a girdle-like structure that secretes mucous during copulation and forms the cocoon.
Term
Copulation
Definition
Sexual intercourse
Term
Cuticle
Definition
a nonliving cuticle secreted by the epidermis covers the body.
Term
Locomotion
Definition
Involves the antagonism of circular and longitudinal muscles.
-when longitudinal muscles contract, the segment swells and setae are protruded
-the previous segment contracts its circular muscles causing the retraction of the setae, pushing forward the anterior segment.
-SEGMENTS MOVE FORWARD AS WAVES OF MUSCULAR CONTRACTIONS MOVE ANTERIORLY
Term
Muscles
Definition
found to the exterior beneath the epidermis
Term
Longitudinal Muscles
Definition
found to the interior of the circular muscles.
Term
Peritoneum
Definition
found anterior to the longitudinal muscles and lining the coelom
Term
Setae
Definition
4 pairs are found on most segments
-two ventrally
-two ventrolaterally that aid in locomotion.
Term
Oligochaetes are SCAVENGERS
Definition
Feed primarily on decaying vegetation (detritus)
Term
Digestive Tract - Oligochaeta
Definition
Food pulled in by Pharynx > Mouth > Esophagus(calciferous glands) > Crop > Muscular Gizzard > Intestine > Anus(and out a nephridiophore)
Term
Pharynx
Definition
pulls food into the mouth and then pushes it down the esophagus.
Term
Calciferous Glands
Definition
evaginations of the wall of the esophagus that rid the body of excess calcium absorbed from food
Term
Crop
Definition
thin-walled food storage structure
Term
Muscular Gizzard
Definition
used for grinding the food
Term
Intestine
Definition
a straight tube where digestion and absorption occurs.
-ends at the Anus.
Term
Typhlosole
Definition
a dorsal unfolding of the intestinal wall that functions in increasing the surface area of the intestine.
Term
Circulatory
Definition
SOMITES 7-11
- expanded segmental vessels referred to as "hearts"
- up to FIVE PAIRS of hearts.
- Main propulsion of the blood is still done by the dorsal and ventral vessels.
Term
Nervous System
Definition
Ventral Nerve Cord & all ganglia have undergone a high degree of fission.
- giant fibers control escape response and small diameter fibers are for locomotion and coordination.
-LACK well-developed eyes and have receptors scattered all over the body surface, especially around the prostomium for MECHANICAL and CHEMICAL reception.
Term
Metanephridia
Definition
the excretory structures
- the funnels of the metanephridia are found in the segment ANTERIOR to the segment containing the tubules and nephridiopore
Term
Chlorogen Tissue
Definition
surrounds the dorsal vessel and over the dorsal surface of the intestine.
**The site of AMINO ACID METABOLISM deaminating amino acids and converting ammonia to urea
- also converts carbs into glycogen and fats
Term
Deamination (fyi)
Definition
Deamination is the process by which amino acids are broken down if there is an excess of protein intake. The amino group is removed from the amino acid and converted to ammonia. The rest of the amino acid is made up of mostly carbon and hydrogen, and is recycled or oxidized for energy.
Term
ALL Oligochaetes - MONOECIOUS
Definition
with cross-fertiliation
Term
Monoecious (fyi)
Definition
(of a plant or invertebrate animal) having both the male and female reproductive organs in the same individual; hermaphrodite.
Term
Cross-Fertilization (fyi)
Definition
Fusion of male and female sex cells from different individuals of the same species. Cross-fertilization is necessary in animal and plant species that have male and female organs on separate individuals.
Term
Testes
Definition
associated with the seminal vesicles.
- SPERM released through the sperm duct openings on the ventral side of SOMITE 15
Term
Eggs
Definition
produced in the ovaries on SOMITE 13
- RELEASED through the oviduct openings on SOMITE 14
- hatch in 1-2 weeks
Term
Seminal Receptacle
Definition
In the grooves between SOMITE 9-10 and 10-11..
- RECEIVE SPERM during copulation.
Term
Seminal Vesicles
Definition
Sites for maturation and storage of sperm PRIOR to their release.
Term
REPRODUCTION OF EARTHWORMS
(Lumbricus terrestris)
Definition
1. Sperm is transferred to the seminal receptacles of each worm.
2. Following coopulation, CLITTELUM forms a cocoon for the deposition of eggs and sperm
-cliittelum secretes a food reserve, albumin, into the cocoon
3. Eggs are deposited into the cocoon through oviduct openings (SOMITE 14)
4. Sperm is released as the cocoon passes over the seminal receptacle openings on (SOMITE 9-11)
5. eggs are fertilized in the cocoon and as the worm continues to back out it deposits the cocoon of fertilized eggs in the soil.
6. Eggs hatch in 1-2 weeks into young worms (NO LARVAL STAGE)that emerge from the cocoon.
Term
Hirudineans are:
Definition
The Leeches
Term
Basic overview
Definition
- MOST are FRESHWATER, but some are marine and some are completely terrestrial, living in warm, wet places.
- more abundant in tropical countries than temperate zones
Term
Feeding
Definition
May be Carniverous or Parasitic
- prey on small invertebrates
- feeding on bodily fluids of vertebrates, sometimes blood
- NOT TRULY PARASITIC because the leech is only on the host for a short amount of time.
- Not species specific, but class specific...
>>>> PREDATORY!!! would be the best term for leeches
- The MOUTH opens in the middle of the anterior sucker on (SOMITE 1)
- may have protrusible proboscis or 3 chitinous jaws
- Secretes Hirudin, an anticoagulent, into the tissue of the host to prevent clotting.
- PHARYNX pumps body fluid from host into esophagus that leads to the stomach with lateral cecae.
- Digestive tract end in a short and anus.
- A leech may go months without food after a feeding.
Term
Body structure
Definition
- DORSOVENTRAL
- adapted for forcing their proboscis into soft tissue
- some have chitinous teeth that can cut through skin
Term
Dorsoventrally
Definition
means it is flattened
Term
Reproduction of leeches - Sexual
Definition
MONOECIOUS w/ a clittelum that only appears during the breeding season.
- Single pair of overies and 4+ testes
- Clitellum on 3 segments in the spring when they breed.
- sperm transferred by penis or by hypodermic injection of spermatophores through the tegment of the other.
- cocoons deposited into soil or attached to underwater objects.
- NO LARVAL STAGE, offspring mature by the following spring.
Term
Monoecious (fyi) and Reproduce Sexually
Definition
(of a plant or invertebrate animal) having both the male and female reproductive organs in the same individual; hermaphrodite.
Term
Number of segments
Definition
EXACTLY 34!!!!
- difficult to extinguish externally
Term
Setae
Definition
reduced or lacking; ABSENT IN MOST
Term
Well-developed Suckers
Definition
at both ends of their bodies
- anterior and posterior
Term
LACK PARAPODIA AND HEAD APPENDAGES
Definition
Term
Annuli (plural)
- annulus (sing.)
Definition
transverse grooves that each segment is marked with externally that give the appearance of secondary divisions in the somite.
Term
Oblique Muscles
Definition
Make Leeches' muscular system more complex than other annelids.
- located between the circular and longitudinal muscles
Term
Hydrostatic cavity
(locomotion)
Definition
used in a looping type of locomotion since the septa are lost.
- connective tissue has invaded the coelom.
- can also swim by undulating their bodies..
Term
Cecae (diverticula)
Definition
a blind tube leading from a cavity or passage.
Term
Gas Exchange
Definition
through the BODY WALL
Term
Highly Modified Coelomic Sinuses (MOST)
Definition
replace the vessels
Term
Coelomic Fluid
Definition
has taken over the function of the blood and respiratory pigments are lacking.
Term
Nervous System - Nerve Ring
Definition
has fused the suprapharyngeal ganglia, circumpharyngeal connectives, and subpharyngeal ganglia into a nerve ring around the pharynx (mouth)
- ventral nerve cords are unfused other than at the segmental ganglia.
- posterior ganglia also form a nerve ring.
Term
Metanephridia
Definition
10-17 pairs of metanepridia...
- one per segment in the middle segments of the leech
Term
Chlorogen Tissue
Definition
found throughout the coelomic cavity
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