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Lab 04
Molluscs
74
Biology
Undergraduate 1
10/05/2006

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Term
2 types of sexual reproduction are:
Definition
monoecious: hermaphrodites (slugs) and sequential herm's (protogyny vs. protandry) Dioecious: sexual selection in heterosexual molluscs. (R-species with many offspring and vs K-species with few large offspring with low mortality)uses hectocotylus
Term
What subclade do tusk shells belong to. Name a synapomorphy of this clade.
Definition
Scaphopoda - has a "spade foot" and lacks gills. Have (and breath through) open ended "tusk"-shaped shell Used for money. [image]
Term
Genus of chiton whose shell is COMPLETELY covered by its mantle.
Definition
Cryptochiton (Gumboot chiton)
Term
One synapomorphy of Bivalva
Definition
2-part shell; no radula but have ciliated siphon to feed and breathe
Term
Why are brown mussels more common the further south you go?
Definition
Brown better reflects the sun = less heat gain.
Term
Operculum- Aperture- What clade would you find these features?
Definition
Operculum is the "door" that closes the aperture (opening). Found in Gastropods[image]
Term
Example of commensalism in clade gastropoda?
Definition
Polychaete worms living with Limpets in their shells.[image]
Term
2 defense mechanisms of Nudibranchs
Definition
Aposematic coloration to warn predators. Eat unfired cnidocytes of cnidaria, move them into their cerrata as predator defense. [image]
Term
Name the 5 daughter clades of Mollusca
Definition
-Polyplacophornas (8 plated chitons), -Bivalves (clams mussels, oysters shipworm etc), -Gastropods (slugs and snails, limpets, nudibranchs) -Cephalopods (octopi, squid nautilus) -Scaphopods (tusk shell)
Term
What do squid use to camouflage?
Definition
chromatophores[image]
Term
How is buoyancy and attitude achieved in cephalopods?
Definition
The chambers in Nautilus act as air compartments with a siphuncle connecting them. Chambers are also present in the "bone" of cuttlefish.
Term
What clade is Tridacna in?
Definition
Bivalva.[image]
Term
What produces/secretes a mollusc's shell?
Definition
Mantle
Term
What is the composition of the 3 layers of a mollusc's shell?
Definition
Outer: proteinaceous layer Middle: calcium carbonate crystal layer Inner: nacreous layer
Term
Which molluscs has 8 shell plates?
Definition
Polyplacophorans
Term
Which mollusc has a symmetrical 2 part shell?
Definition
Bivalve
Term
Only ________ __________ retain the (coiled) shell.
Definition
Ancestral cephalopods
Term
Describe early mollusc development and how it moves at that stage.
Definition
Trochophore larvae that move w/ cilia band in midregion.
Term
How are molluscs and annelids related?
Definition
Development: protosomes, trocophore larval stage, Anatomy: body cavity lined with mesoderm
Term
Name the 2 body regions molluscs have and their functions.
Definition
1. Muscular foot: mostly for locomotion 2. Dorsal visceral mass: contains internal organs and covered with mantle[image]
Term
What does the mantle cavity contain in a mollusc?
Definition
Gills, nephridopore, and anus
Term
What factor can make a mollusc's shell colorful?
Definition
Mineral content in middle layer of shell can make the mollusc colorful.
Term
How do pearls form?
Definition
A mollusc's mantle secretes a nacreous layer around a foreign object.
Term
What structures are found on the muscular foot of a mollusc?
Definition
The head with a mouth, and sensory organs are found on the anterior portion of the muscular foot.
Term
What is the unique structure found in a mollusc's mouth?
Definition
A radula: a ribbon-like, belt-shaped, raspy tounge.
Term
How does radula structure reflect diet?
Definition
Herbivores have many small teeth. Carnivore have fewer and larger teeth.[image]
Term
How are Chiton radulas unique?
Definition
Contain iron oxide, since they lick algae off rocks.[image]
Term
What does this photo show?[image]
Definition
A moon snail and it prey (clam)
Term
How are cone snails' radulas specialized?
Definition
Cone snails havs a ciliated siphon in its mantle that is chemical senistive, and modified radula that is harpoon-like to inject calicum channel blocking neurotoxins into their prey.[image]
Term
How do mussels in rivers increase increase reproductive sucess?
Definition
Mussels in rivers use a minnow-like mantle margin to attract fish. When the arrives, the mussel releases its offspring which parasitize the fishes gills. The glochidia recieves protection and is dispersed further than otherwise by this parasitism. [image]
Term
How do river mussels's offspring hold onto their host fish, and what benefit do they receive from the fish?
Definition
With teeth that bite into gill filaments by contracted adductor muscles. The offspring recieve protection and upstream transport. [image]
Term
How is Darwin's concept of sexual selection observed in male dioecious cephlapods?
Definition
Many male cephlapods have hectocotylus arm that collects sperm from its reproductive tract until copulation. Upon copulation arm detaches in the female's reproductive opening, blocking other males from reproducing with her.
Term
Define protogyny.
Definition
Starting reproductive life as a female. When the organism reaches a size in which it can compete with males sucessfully, the gender changes from female to male.
Term
Define protandry.
Definition
When an organism starts its mature life as male, then becomes a female is an example of protandry.
Term
What is the benefit protogyny?
Definition
An organism can be producing offspring throughout its lifetime, with male producing more offspring due to the greater numbers of fertilization events in which they can participate in.
Term
What is the benefit of protandry?
Definition
If there's no male-male competition, then being big as a female = leaving more young in lifetime.
Term
Which mollusc subclade has a spade foot?
Definition
Scaphopoda
Term
How do Scaphopods feed?
Definition
Burrow spade foot into the soft ocean bottom and feed on detritus caught in their tentacles.
Term
How do scapohods respire?
Definition
Beating cilia to drive water in mantle cavity through the holes at each end of their shell. Tusk shell organism lack gills.
Term
What mollusc was used as money by Native Americans?
Definition
Scaphopods -- tusk shells
Term
What mollusc has >2 segments, how many, and why?
Definition
Polyplacophorans have 8 jointed shell segments which allow them to conform to irregular intertidal surfaces.[image]
Term
What do chitons eat?
Definition
Algae
Term
What is a home scar?
Definition
A home scar refers to specfic depression in a rock a chiton or limpet fits into.
Term
How do chiton respire?
Definition
By beating cilia through their mantle cavity, across a pair of gills each located in one of 2 lateral mantle cavity grooves. Ex: gumboot chiton.[image]
Term
What is unique to the radula of chitons?
Definition
Iron in largest teeth aids in licking algae off rocks.
Term
How do chitons reduce predation?
Definition
Extending beyond their shell is mantle which is often covered with defensive bristles.[image]
Term
Name 2 sister clades which have lost their cephalization?
Definition
Bivalva and scaphopoda have lost their radula.
Term
How do bivalves respire and eat?
Definition
Bivalves respire via ciliated incurrent and excurrent siphons which pumps oxygen and food rich water into their shell. Food is filterd by palps.
Term
Why are the adductor muscle scars on a clam so obvious?
Definition
The adductor muscle scars are so obvious because this was the location at which the adductor muscles attached to the clam shell with the purpose of opening and closing the clam's shell. [image]
Term
Are older or more recent fossils more accurately carbon-dated? What does the graph look like?
Definition
More recent fossils are: as the ratio of C14 to C12 gets closer to zero, so little C14 remains that the margin for error increases signifigantly.[image]
Term
Give some synapomorphies of clade Mollusca
Definition
1) start off as trochophore larvae w/ a band of cilia for movement.

2) 3-layer shell secreted by

3) mantle
Term
What are ocelli?
Definition
Simple eyes found on scallops which watch for predators.[image]
Term
What is the typical environment for a mussel and how are they anchored to that environment?
Definition
Rocky intertidal (and subtidal), attaching with byssal therads secreted by a gland in its foot.
Term
How do non-sessile species of bivalves, escape predators?
Definition
Rapid contraction of adductor muscles allow for jet propulsion in bivalves to escape predators.
Term
What clade and genus is this organism:[image] What is their greatest predator?
Definition
Clade Bivalve, genus Tridacna (giant clam).
Humans are the giant clam's greatest predator.
Term
What are adapations of burrowing bivalves which live in a hard medium?
Definition
Shipworms: bores into rock or wood, & have reduced shells usually used for abraiding their inhabited surface Sandstone borer: bores into sandstone with a rough posterior shell and chemical secretions. Here is a picture of a sandstone borer.[image]
Term
What are adaptations of bivalves which inhabit soft mediums?
Definition
1) well developed shells for protection 2) long and thin siphons for feeding, breathing and waste removal due to their immobility. This is an example of a long siphoned clam.[image]
Term
Scientific name of chiton whose mantle
PARTIALLY grows over shell.
Definition
Katherina
Term
90% of molluscs are in which subclade?
Definition
Gastropoda - "belly-footed" snails and their allies.
Term
What's the gastropod's "opening" & "door" called.
Definition
aperture = opening operculum = door[image]
Term
Name the molluscs that have a shell that rotates into a coil.
Definition
Gastropoda & ancestral Cephalopoda (a synapomorphy they share)
Term
Name the developmental characteric of gastropods involving rotation.
Definition
Torsion. The mantle cavity is posterior in embryos, then the viseral mass rotates 180° bringing the mantle cavity above their head.
Term
Slugs undergo uncoiling. What is this?
Definition
Detorsion. An evolutionay reversal of the 180° rotation of their immediate ancestors.
Term
The limpet clade is ...? What's unusual about their shell?
Definition
Limpets are Gastropoda. Their shell ISN'T coiled, but cone shaped. [image]
Term
Name this mollusc, its clade and a key characteristic to reduce competition with other organisms in its habitat. [image]
Definition
Owl limpet. Gastropoda. The design is compromised of prying ability vs. hydrodynamics. A squat offset shape allows it to bulldoze competitors out of its habitat for better access to food.
Term
Name the clade, name, and a unique morpological characteristic for defense of this mollusc:[image]
Definition
Gastropoda. Keyhole limpet. This gastropod can raise its girdle upward to keep starfish from eating it.
Term
Which clade of molluscs have diverse colors and patterns in tropical regions and why?
Definition
Gastropoda. Disruptive coloration can aid in camoflaging in tropical areas of high predator density
Term
Name the clade of this gastropod which glues other shells to itself and why it does this.[image]
Definition
Gastropoda (decorator snail). This gastropod glues shell to itself to look different and to make it more difficult to swallow.
Term
Slugs and what lack shells in what clade?[image]
Definition
Nudibrnach , gastropoda
Term
What is the clade and name of this mollusc?[image]
Definition
Gastrpoda, sea hare (Aplysia)
Term
What do nudibranches prey on, what is sequestered in projections on their back, and what purpose does it serve?
Definition
Nudibranches prey on cnidarians which carry nematocysts. The nematocysts are then sequestered in cerrata on the nudibranches back. This serves as aposematic coloration to warn predatory fish aganist eating them.
Term
Did nudibranches evolve aposematic coloration to benefit the fish that prey on them?
Definition
No. This trait serves the nudibranch to enhance its survival chance, and was selected for.
Term
Name the clade, name and unique features and adaptations of this animal:[image]
Definition
Cephlpoda. Squid. Highly developed eyes, large brain, keen predator, 3 hearts, beak-like jaws,2 long tentacles with sucker pads (with teeth in larger species), skin pigment varitability due to chromatophores,
Term
Where do eyes arise from?
Definition
A common ancestor: eyes evolved once with the pax-6 gene.
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