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Evolution
N/A
322
Biology
Undergraduate 3
11/14/2012

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Term
Why does undergoing sexual selection put organisms at a disadvantage if the environment never changes?
Definition
Because the organism will have to devote time and energy into finding a mate to reproduce with when it is not necessary. An organism that undergoes asexual reproduction just clones itself so it only takes as much time and time and energy as it takes to produce another like itself. Assuming the organism is well adapted to the environment asexual reproduction will increase the population numbers more effectively.
Term
Why is sexual reproduction at a disadvantage in unchanging environments?
Definition
1) it takes more time and energy to find a mate and impress it than it takes to just undergo asexual reproduction.
2) Asexual reproduction will increase an organism's numbers more effectively assuming the organism is adapted to the environment.
Term
Why is sexual reproduction a good thing in most cases?
Definition
It causes variation and breaks up linkage disequilibrium.
Term
What is linkage disequilibrium.
Definition
Linkage disequilibrium is when two alleles are linked on the same chromosome.
Term
What is sexual dimorphism?
Definition
It is where the male and female of a species look different.
Term
In sexual dimorphism what sex will almost always become elaborately decorated? Why?
Definition
The male will always be more showy than the female because the females are almost always the ones selecting mates.
Term
Why is female selecting males for certain traits selected for by natural selection?
Definition
Because females select males with large antlers or colorful plumages. The production of these excesses means that the male is fit and will pass on quality genes to the offspring. In the case of antlers and other similar structures, the females might be selecting males that are best at defending their teritory, thus proving their fitness.
Term
Why are females the ones that select their mates?
Definition
Because there is little metabolic investment involved in producing sperm, but a huge amount of investment involved in producing eggs. the female also has to go through pregnancy and usually raise the offspring in the few cases where the offspring is raised. often times males play no role in care for the offspring. Females can not produce an unlimited number of offspring. All of this means that females only have a couple of chances to produce viable offspring to pass on the mother's genes.
Term
Which gender is MUCH more likely to not produce any offspring at all?
Definition
Males (majority are unsuccessful in some cases)
Term
Does having multiple partner increase reproductive success more in males or females?
Definition
Males.
Term
Why do we find that females from the species Sygnaphus typhle (broad nosed pipefish)often die without reproducing where males almost always have reproductive success?
Definition
Because the males in Sygnathus typhle have to carry the developing embryo in a sac on their ventral side. This means the female only invests an egg, though expensive metabolically, she does not have to bear the offspring.
Term
What is intrasexual selection? What causes it?
Definition
Intrasexual selection is when members of the same gender with low metabolic investment in sexual reproduction (usually males) compete for the opportunity to mate. It is caused by sexual selection for the most fit individuals by the sex that has the highest metabolic cost in reproduction.
Term
What will be the result of territory defense intrasexual selection?
Definition
Increased size of the defense structure. (horns for example)
Term
Why dont the horns of a ram grow indefinitely?
Definition
because the metabolic cost of creating horns that are too big will mean more food will need to be consumed to sustain them. This means the size will be stabilized at some intermediate size.
Term
Why do males often defend a territory from other males?
Definition
Because they don't have to invest much in reproduction, they adopt strategies to increase their reproductive success (since i males reproductive success is determined by the amount of mates he can sucessfuly breed with)
Term
Where will the most fit males be in populations that have intra-sexual selection?
Definition
in the highest quality territory. More females visit the higher quality teritory so it is more coveted.
Term
sexual selection leads to....
Definition
Sexual dimorphism
Term
How do the coho salmon reproduce?
Definition
The females deposit eggs on the bottom of the river bed. The older hooknose males fight for a position in line to ejactulate on the eggs. They end up lining up according to size. The smaller Jacks don't stand a chance so they evolved a more sneaky approach to compete. They hide nearby while the female coho salmon is laying her eggs, then quickly swims over and ejaculates on the eggs.
Term
How does the Mediterranean fruit fly deal with the fact that competition for females is fierce? (females mate with multiple males)
Definition
They deposit 2 times as much sperm in the presence of another male.
Term
What is the correct term for a sperm delivery organ that does not also function as an excretory organ?
Definition
Aedeagus.
Term
What adaptation have damselfly and dragonfly species males developed to deal with competition from other males to mate with females? What is the counter adaptation?
Definition
The males deposit a cement-like substance after they copulate to plug up the cloaca. Males have counter adapted hooks at the end of their aedeagus to scrape out the plug and at least some of the sperm.
Term
What are the groups called that lions live in?
Definition
Prides.
Term
What gender of lions is born in the pride and stays in the pride, and what gender is unrelated to the pride?
Definition
The females are born into the pride and stay in the pride. The males come from outside to prevent imbreeding.
Term
What is it like being a male lion in a pride?
Definition
The male lion leaves the pride he is born into and joins another pride. He must then drive out the male that is currently mating with the females. The female lions can not become pregnant until the cubs are fully weaned. To solve this problem, male lions simply kill off all the cubs from the male that they out-competed. This will bring on fertility faster.
Term
Why are red collared willow birds much larger than the females? Why do they have such enormous tails?
Definition
Sexual dimorphism = Sexual selection.
Term
Why does the tree frog inflate it's throat?
Definition
To create a resonance chamber for it's mating call.
Term
Which male tree frog will be chosen by females?
Definition
The one who has the longest mating call. Other males will also be calling, the males must increase the length of their call to distinguish them from the others.
Term
what is the phototaxis score in the experiment testing female tree frog's response to male calls?
Definition
The phototaxis score is the ratio of the time it takes a female to approach a control divided by the time it takes a female to approach the experimental.
Term
How did scientists figure out that female tree frogs prefer longer male calls?
Definition
They set up a control and started measuring the amount of time it takes a female to approach a loudspeaker playing different length calls. They then divided the time it took the control divided by the time it took the experimental to find the phototaxis score for the Y axis of a graph and they placed the length of the call on the X axis. The graph shows that females strongly favor males with longer calls.
Term
Alison welsch wanted to carry out an experiment to test if tree frogs have to wait until they are of reproductive age to reap the benefits of having a father with a long mating call. How did she do this? What were the results?
Definition
She took eggs from random females and mated one set with sperm from a short calling frog and one set with sperm from a long calling frog. She then divided the offspring into two groups, one group of LC/SC with ample food, and one group of LC/SC was given a low amount of food. They observed the fitness. They measured time it took to get to metamorphosis, mass of the tadpoles, and the rate of growth after metamorphosis. The results were mixed but always showed that LC were better at surviving.
Term
What does the male rattlebox moth offer the female rattlebox moth for mating?
Definition
Life insurance. The male deposits a chemical defense on the females after mating that causes spiders to release the moths caught in their web.
Term
what does the male bittacus epicalis (hanging fly) offer the female for mating? What males will be selected for?
Definition
A blowfly that the male uses to distract her long enough for copulation to take place. She disengages when she is finished eating. Males who bring bigger blowflies will be selected for, up to about 20 minutes. If the female is not done with the blowfly after 20 minutes, the males take the blowfly back and use it on other females.
Term
traits that females prefer in mates will _____ in offspring
Definition
Increase the fitness
Term
Females are more choosey with the mates they choose if ____
Definition
There is a high investment in the offspring for the female.
Term
T/F In most case females are the ones who are choosey.
Definition
True
Term
can water mites see?
Definition
Not very well.
Term
How do water mites eat?
Definition
They can sense the vibrations that copepods make then they wave there arms around looking for it.
Term
How do water mites mate?
Definition
The male can smell the female, but the female might have left, so he must make sure she is still there. He waves his legs at the frequency of a water mite. The female lunges towards him which proves she is there. The male then drops a spermatophore (sperm packet) on the leaf, and vibrates his arms again. This time the female is fertilized.
Term
what are two theories that explain which how the trembling behavior and the netting behavior of water mites arose?
Definition
1) the trembling arm behavior arose, then the feeding behavior arose, then the trembling behavior was lost in one species.
2)the netting behavior arose, then the trembling arose on two seperate occasios.
Term
what is the sexy sons hyothesis?
Definition
Fathers with a trait that females prefer will pass it on to their offspring who will be more attractive to females like their mother. The trait does not necessarily improve fitness, it is just what the female finds appealing.
Term
why do spotted cucumber beetles adhere to the sexy sons hypothesis?
Definition
The male that strokes the females antennae fastest are selected by the female for copulation. The only increase in fitness is due to the fact that females like fast stroking cucumber beetles.
Term
What is the sexual dimorphy in diopsids (stalk eyed flies)?
Definition
Males are much larger than females and have longer eye stalks.
Term
If the male diopsid offspring has long eyestalks, the female offspring will...
Definition
Prefer longer eyestalks in a mate.
Term
How do they know that diopsids who were born from short eyestalk parents prefer short eyestalk mates?
Definition
They did an experiment where they raised one group with only short eyestalks, one with only long eyestalks, and a control. They then let each group choose mates from both short and long eyestalked mates and the short eyestalk group chose all short eyestalked mates, the long eyestalked group chose all long eyestalked mates.
Term
are eyestalk length and preferance heritable?
Definition
Yes.
Term
can selection on one trait produce a response in another trait?
Definition
Yes.
Term
Why do diopsids always choose long eye stalked males in nature?
Definition
Because the long stalked males are more fit only because they have Y-chromosome allele that produces more sons than daughters. Females outnumber males. Females that mate with long eyestalked mates are more likely to produce more sons.
Term
What is polyandry?
Definition
Having multiple mates.
Term
How do you run a paternity test?
Definition
DNA from the mother, father, and baby are all collected and placed on an agarose gel. The gel is then placed into radioactive DNA solution, this binds to specific DNA sequences on the agarose gel. they run the samples on the gel and if the baby's dna lines line up with ones seen in the mother and father, they are the parents. If not, the one that the baby does not line up with is not the parent.
Term
Why do female blackbirds nest with one bird but also go out to be fertilized by other birds?
Definition
She wants to make sure all of her eggs are fertilized. This will increase her reproductive success.
Term
Why do prairie dogs have multiple partners?
Definition
To increase the success rate of reproduction. They increase the size of their litter this way and make sure they produce offspring.
Term
what do male pipefish choose in females?
Definition
Larger females with larger skin folds and a lower number of black dots due to parasites.
Term
Can sexual selection select for certain behaviors?
Definition
Yes
Term
What are phylogenies used for?
Definition
to create hypotheses for the evolution of a species.
Term
Do plants often show sexual dimorphism?
Definition
Yes.
Term
Do plants often show sexual dimorphism?
Definition
Yes
Term
In plants, which are usually bigger, male flowers or female flowers?
Definition
Male flowers.
Term
What is the phenotypic result of sexual selection in male flowers?
Definition
sexual selection causes more showy flowers.
Term
Explain the culture of the Yanomamo people?
Definition
They live in the jungles of South America and are very war like. They raid and kill the populations of rival villages. 40% of males have had some involvement in killing someone else. They earn the title "unokai"
Term
Do women favor the Unokai (killers) in the Yanomamo society?
Definition
Yes, Unokai have more wives and more children on average.
Term
Why is it hard to study sexual selection in humans?
Definition
Because attractiveness to certain traits varies based on what culture you are raised in.
Term
Why is donald trump like a hanging fly?
Definition
He gets a lot of women because he has money. Money is a resource just like the blowfly the hanging fly uses to distract while mating.
Term
One study shows that women usually select men who are...
Definition
Tall.
Term
What is usually attractive to the opposite sex in humans?
Definition
Indicators of good health.
Term
Whales are the cousins of...
Definition
Hippos
Term
What are the two main objectives of evolutionary biology?
Definition
1. to understand the process of evoluion
2. to recover the history of life on earth (systematics)
Term
What is the difference between taxonomy and systematics?
Definition
Systematics is interested in grouping organisms into groupings based on their evolutionary relationships. Taxonomy assigns organisms into groups usually based on common characteristics.
Term
What are sister taxa?
Definition
Two groups that are closely related to eachother through a recent common ancestor.
Term
What is a monophyletic group?
Definition
A group containing the common ancestor and all it's decendents.
Term
Do paraphyletic groupings reflect evolutionary history?
Definition
No.
Term
List an explain the 4 main sources of data for systemic studies.
Definition
Morphology - the physical appearance and physical traits the organism has
Behavioral - the actions of an organism
Ecologial - the place the organism lives, breathes, and feeds.
Molecular - genetic data in the form of gene order, chromosomal inversions, protein or DNA sequences, etc.
Term
What are the objectives of systematic biology?
Definition
1. to resolve unknown relationships
2. to test previously advanced hypotheses
3. to trace the evolution of a particuar character or set of characters.
4. develop new methods for analyzing data
Term
Taxonomy can result in ______ groups.
Definition
Paraphyletic
Term
can taxonomy answer or ask evolutionary questions?
Definition
No.
Term
What is taxonomy?
Definition
The science of identification and classification.
Term
what is the problem with the appeal to authority way of approaching evolution?
Definition
Who is to say who is the most authoritative on the issue? Not all experts agree. The results are not repeatable.
Term
What is another name for numerical taxonomy?
Definition
Phenetics
Term
What is Phenetics? What does it not take into account?
Definition
Phenetics is the grouping of organisms based on their overall similarity. Systemics does not take convergent evolution into account.
Term
What is convergent evolution?
Definition
The independent acquisition of similar charactersin different evolutionary lineages.
Term
give an example of convergent evolution
Definition
Bicameral eyes, wings.
Term
What is parallel evolution? Why is it different then convergent evolution?
Definition
Parallel evolution is the independent acquisition of a similar character in closely related lineages. In convergent evolution, they are not closely related lineages.
Term
What are reversals?
Definition
A reversal happens when a mutation returns a character to it's ancestral condition.
Term
What is homoplasy? What is it's opposite?
Definition
Homoplasy is a catchall term for convergent evolution, parallel evolution, and reversals. Opposite is homology.
Term
Why does homoplasy confound systematic studies?
Definition
Because it incorrectly implies common ancestry.
Term
What is homology? give an example.
Definition
Homology is similarity in traits due to a common ancestor. An example would be the bones in the arm of every mammal.
Term
Monophyletic groups represent evolutionary history T/F
Definition
true
Term
Paraphyletic groups represent evolutionary history T/F
Definition
False
Term
What are phylogenies?
Definition
Evolutionary trees, they reveal the pattern of evolution.
Term
What is phylogenetics concerning?
Definition
Evolutionary relationships
Term
Phylogenetics seeks to organize classifications into...
Definition
monophyletic groups.
Term
Who was the first person to suggest that evolutionary relationships could be organized into a tree-like diagram?
Definition
Charles Darwin
Term
When was the age of numerical taxonomy?
Definition
the 1950's
Term
Who is Willi Hennig?
Definition
The father of cladistics. A german entomologist
Term
Does cladistics take into account both homology and homoplasy?
Definition
Yes
Term
What is an ancestral character called?
Definition
A plesiomorphy
Term
What are derived characters called?
Definition
Apomorphies
Term
With respect to the evolution of eukaryotes, is the chloroplast a plesiomorphic or apomorphic character?
Definition
Apomorphic
Term
With respect to the phylogeny of the plant, is the chloroplast apomorphic or plesiomorphic? Why?
Definition
Plesiomorphic. The chloroplast was in the ancestor of all plants. We know this because all plants have seeds.
Term
when can you say that seeds are an plesiomorphic character?
Definition
If you are looking that the phylogeny of seed plants.
Term
When did land plants first evolve?
Definition
500 million years ago.
Term
What are bryophytes? What evolutionary significance do they have?
Definition
Bryophytes are mosses, they are thought to be the oldest existing lineage of plants.
Term
In what plants does a vascular system first evolve?
Definition
Pteridiophytes AKA Ferns.
Term
what characters evolve in plants?
Definition
Mosses - Vascular system - Seeds - Flowers
Term
Using synapomorphies, synplesiomorphies, apomorphies, and plesiomorphies is all part of _______ method.
Definition
Willi Henig's Cladistics
Term
What is the term for shared ancestral characters?
Definition
Synplesiomorphies
Term
What is the term for shared derived characters?
Definition
Synapomorphies.
Term
Homology is evidence of...
Definition
Divergent evolution
Term
Why is cladistics based on synapomorphies and not synplesiomorphies?
Definition
Since all members have the synplesiomorphic trait, it does not help us group organisms.
Term
What is Hennig's grouping rule?
Definition
Synapomorphies are evidence of common ancestry. Synplesiomorphies, convergences, and parallelisms are not evidence of common ancestry.
Term
What is Hennig's Auxillary Principle?
Definition
Never assume convergent evolution or parallel evolution, always assume homoplasy in the absence of contrary evidence.
Term
How do you create a cladogram?
Definition
You look at various different orgnisms and figure out whether they have a trait or don't. arange them accordingly assuming homology.
Term
What is the point of making an unrooted phylogenetic tree?
Definition
They depict relationships but not the order of divergence. They simply group sister taxa.
Term
What animal is thought to be the ancestor of all birds?
Definition
Archaeopteryx
Term
Can the outgroup be a member of the group you are studying?
Definition
No.
Term
Where on the cladogram is the outgroup?
Definition
at the very begining.
Term
What do you call the group you are studying?
Definition
The ingroup.
Term
What charactaristics must your outgroup have?
Definition
It must have some characteristics similar to your ingroup.
Term
What is the purpose of the outgroup?
Definition
To help you decide the order that the taxa diverged. The taxa that are most like the outgroup arose first.
Term
What is the term for a derivedcharacter that is not shared?
Definition
Autapomorphy
Term
What is a reversal?
Definition
When a derived species goes reverses to an ancestral characteristic instead of a derived one.
Term
What is occam's razor?
Definition
The simplest explanation is always the best.
Term
How can we use the axillary principle to solve a problem where we have to choose between multiple trees?
Definition
We choose the tree that has the least amount of homoplasy.
Term
What is the principle of parsimony?
Definition
The answer that requires the fewest changes is preferred to more complex answers.
Term
How can you apply the principle of parsimony and Occam's razor to a phylogenetic tree?
Definition
If you have conflicting trees you should choose the one with the least amount of evolutionary steps.
Term
What is a consensus tree?
Definition
a consensus tree is a tree you make when you have homoplasy and end up with two or more trees that are equally plausable. The idea is that you colapse them intoone tree called a polytomy with all taxa coming from one node.
Term
Which cladogram contains an outgroup: rooted or unrooted?
Definition
Rooted
Term
What rule implies that the cladogram with the least number of steps is prefered?
Definition
Both the principle of parsimony and Occam's razor.
Term
Both cladistics and Phenetics is preferable to...
Definition
Appeal to authority.
Term
The taxa in a phylogeny involve the _____ of different organisms.
Definition
Taxonomy
Term
Cephalopod mollusks and vertebrate both have a..... this is an example of....
Definition
Camera eye
Convergent evolution.
Term
Why do systematists use a computer program to erect phylogenetic trees?
Definition
Because phylogeny actually involve a complex set of mathematical equations called NP completeness problems. There are so many possibilites that it would be impossible to sort through the possibilities.
Term
Is it more of a problem in systematics to add a character or a taxa?
Definition
A taxa because there are many more possibilities.
Term
The fastest computer programs don't have enough time to check all the possibilities in _______ taxa or more.
Definition
12
Term
Beyond 12 taxa _________ is employed which gives an approximate best result.
Definition
Heuristic search method
Term
How does the computer program that searches different phylogenic trees work?
Definition
It makes a random tree using the information you gave it then it rolls through the possibilities. Each time it checks the number of changes, and if it's lower than the previous it throws the previous out.
Term
What are the Chiropterans?
Definition
The Bat family
Term
What group are the primates most closely related to?
Definition
Chiropterans. Bats!
Term
What are perissodactyls?
Definition
Mammals that have hooves and an odd number of toes such as horses and Rhinos
Term
What are the Artiodactyls?
Definition
Even toed ungulates. Hippos, pigs, deer, cows.
Term
Where did they used to think the hippo belonged on the cladogram?
Definition
As a sistergroup to the hippos.
Term
Explain how the Whippo hypothesis came about.
Definition
John Gatesy sequenced a bunch of milk proteins and used them to build a phylogenetic tree. That tree showed the whale was most closely related to the hippo.
Term
What debate did the whippo hypothesis spark? Between what two groups of evolutionary biologists?
Definition
It sparked a debate in the scientific community on the imporance of molecular evidence. The morphologists said that morphological differences are based on many different gene interactions, and the molecular systemists said that DNA base pair analysis reduces the amount of homoplasy.
Term
What chalenges did gatesy face with the whippo hypothesis?
Definition
People said that since he used the heuristic search method he may not have got the most parsimonious tree.
People used to astragalus to raise the point that it is not a artiodactyl like astragalus, this would give the tree 2 steps, non-parsimonious.
Term
What evidence led to more acceptance of the whippo hypothesis?
Definition
Nikaido used LINE's and SINE's to construct a phylogeny that also puts the whale as a sister taxa to the hippo. Finally widespread acceptance came when fossils were found of transition states between the hippo and the whale with the inner ear bones unique to whales, and the astragalus of the artiodactyls.
Term
What is a LINE?
Definition
Long Interspersed Elements. Genome insertions of mRNA reverse transcriptase.
Term
What is a SINE?
Definition
Short Interspersed Elements. They are similar to pseudogenes and they are thought to arise by reverse transcriptase of mRNA.
Term
What are sources for molecular data?
Definition
Presence or absence of genes
Inversions
Gene duplications
Chromosomal inversions or translocations
Microsatellites
Haplotypes
Gene and nucleotide sequences
Protein sequences
Term
What is FOXP and what is the only extant species that has it?
Definition
FOXP is the gene that gives humans the ability to speak. Only humans have it. Neandertals also had it.
Term
If you find that two species have a chromosomal inversion at the same spot, what does this suggest?
Definition
The inversion happened in the common ancestor to these two animals.
Term
How do microsattelites help identify synapomorphies among taxa?
Definition
the presence of microsatellites in the same spot proves linkage between two taxa.
Term
What are the advantages of using gene sequences to create a phylogenic tree?
Definition
You have use many different characters. This allows you to avoid polytomies and get a more accurate tree becase you have the same number of species and more characters specific to that species. A single gene or protein contains hundreds or even thousands of different characters because every amino acid or nucleotide base is a character.
Term
When do you get polytomies?
Definition
When you don't have enough variation in your characters to tell which evolved first.
Term
a tree that has no polytomy is ______ compared to a tree that has polytomy
Definition
Resolved, more resolved.
Term
Why is it easier to recognize synapomorphy using molecular analysis with genes and proteins?
Definition
There ar only 20 different kinds of amino acids and 4 different nucleotides which means there is a limited number of character states. This makes it easier to recognize homology.
Term
What is the disadvantage of gene and protein analysis for systematics?
Definition
There are only 4 different nucleotides and 20 different amino acids, so it is harder to recognize homeoplasy. Basically, when looking at a gene sequence, many different mutations may have taken place and they both may have converged on nearly identical gene sequences.
Term
How do you compare two gene seqences?
Definition
First you identify homologous sequences
Second you align the sequences with one another so you can see which nucleotides are the same and which differ.
Third you identify substitutions, deletions, and insertions
Fourth you add gaps where insertions/deletions shifted the genome.
Term
Why do you add gaps to the gene sequences you are comparing?
Definition
Deletions and insertions shift the whole genome. Adding gaps shows the deletion but also realigns the genome to show similarities again.
Term
Why is adding gaps hard?
Definition
Sometimes you are left with more than one possibility and you can't be sure which one is accurate.
Term
How do computer algorythms solve the problem of when to add gaps to the gene sequence?
Definition
It uses program settings such as the gap to change cost - basically it is programmed to only put a gap in if there is more than two nucleotides misaligned in a row.
Term
What do you do once you have properly aligned the gene sequences?
Definition
You put it into a phylogenic tree program to try to find the best fit in the tree.
Term
What are the advantages of parsimony?
Definition
All characters are considered independently - there is no loss of character information.
It is the best method for combining different sources of data.
It makes the fewest assumptions.
Term
What are some objections to the parsimony method?
Definition
It can ignore the idea that DNA similarities in base pairs are due to convergent evolution. They assume that everything is due to homology as long as there's no evidence to the contrary. The "long branch" problem. Basically, organisms that split from a common ancestor a long time ago are more likely to undergo convergent evolution. Cladistics does not have any statistical values that allow scientists to compare it to other hypotheses. "the best tree is the shortest tree" means that if you get a short tree with some very suprising relationships, and a slightly longer tree will have more traditional relationships, it's hard to argue that the shorter one is better.
Term
What is a long branch attraction probelem?
Definition
The more distantly related 2 species are the more likely they are to undergo convergent evolution. This tricks cladists into thinking species distantly related are actually very closely related.
Term
Some systemetists do not like using cladistics, what do they use instead? Which method is most commonly used as an alternative?
Definition
Distance methods.
Statistical methods like maximum likelihood.
Maximum likelihood is more popular.
Term
What are primary homology statements
Definition
The alignment of the Gene sequences.
Term
What is the distance method?
Definition
Basically it is phenetics applied to molecular data. Similar sequences are placed together as sister taxa on a tree. The amount of difference in the nucleotides are used to estimate how distantly related two organisms are.
Term
What are some benefits to the distance method?
Definition
It is the fastest method of analysis and it always produces a fully resolved tree.
Term
do most systemists use distance methods? are they skeptical about distance method results?
Definition
No. yes.
Term
what is bad about the distance method?
Definition
an entire gene sequence of information is reduced to one number. There is no character by character analysis like in other methods.
Term
Why is maximum likelihood used more today than it was 10 year ago?
Definition
Computing power has increased enough to allow us to perform the complecated calculations involved.
Term
What happens in the Maximum likelihood method (statistical methods)?
Definition
You enter your data into the computer then the computer will analyze it and tell you what program you should use. It assembles the data into phylogenetic trees by doing something similar to the heuristic method - searching for shorter and shorter trees. Then it calculates the likelihood ratio of each tree, the lowest likelihood ratio is selected as the one that best fits the data.
Term
What are some advantages of maximum likelihood?
Definition
It uses each individual character. There is no loss of character info.
Can determine if a specific hypothesis is better than another because it caculates likelihood.
Term
What are some problems with the maximum likelihood method?
Definition
Your results depend on your model. If you do not choose the right model paralleling how evolution took place in this case, your results will be meaningless. However, the models are estimated from the data.
You can't combine models. If you want to combine molecular and morphological data, you will have to come up with a model for how the morphological characters evolved.
Maximum likelihood is computationally intensive.
Term
What are the three molecular evolution models?
Definition
1) Jukes Cantor - all mutations are equally likely
2)Kimura 2 peramiter model - unequal rates of trasitions and transversions
3)General Time Reversable model (GTR) - a different rate for each possible change.
Term
What is good about the Jukes Cantor model?
Definition
It doesn't make any assumptions there's no evidence for.
Term
What is a transition ?
Definition
A purine (A or G) is turned into another purine. Or when a Pyrimidine is turned into another Pyrimidine (C T)
Term
What is a transversion?
Definition
A pyrimidine (C or T) is turned into another pyrimidine.
Term
Purines
Definition
A G
Term
Pyrimidines
Definition
C T
Term
What did Kimura find out about molecular biology?
Definition
Transitions occur more frequently than inversions.
Term
What are the specifics of Kimura's model?
Definition
Transitions are given a value of 1 and transversions are given a value of .5
Term
Why do they think the reason is that transitions happen more than transversions?
Definition
The shapes of bases prefer to be replaced by similarly shaped bases.
Term
Why is the General time reversible model sometimes used?
Definition
Because sometimes the rate is different for each type of transition/transversion.
Term
Is the most complex model always the best model to use?
Definition
No. The best model to use will be calculated by the Maximum Likelihood program.
Term
Which will be more variable in a gamma distribution: More variable sites or less?
Definition
less variable sites.
Term
What is the difference between bootstrapping and jacknifing?
Definition
Bootstraping means the machine will pull out a random sample of your characters, replace it, and repeat pulling out samples and testing then replacing them. Jacknifing, on the other hand, does not replace the characters it tests.
Term
What is a bremer support value?
Definition
The amount of homoplasy that would be necessary to colapse your set of relationships into a polytomy
Term
What is horizontal transfer?
Definition
When bacteria exchange DNA with each other.
Term
Which two types of organisms can transfer dna from themselves to another organism?
Definition
Viruses and bacteria.
Term
What is an example of a way that there can be no ancestor to decendent relationship in the genes?
Definition
Horizontal transfer
Term
How can you avoid the problem of possibly tracing a gene which may have undergone gene transfer instead of tracing the organism?
Definition
Analyze multiple genes.
Term
Why do systemetists analyze multiple genes?
Definition
Because one gene may have undergone horizontal gene transfer, you want a large sample of genes that give the same results.
Term
What must you choose carefully when running a gene analysis to erect a phylogeny?
Definition
The gene regions you use. You don't want to use an ancestral gene region because that would be useless. No variation. This would be like using a chloroplast to erect the plant phylogeny.
Term
do different gene regions acquire mutations at the same or different rates?
Definition
Gene regions acquire mutations at different rates.
Term
What mutation rate do you want in a gene?
Definition
You want a mutation rate that is fast enough that you can recognize variation across species, but slow enough that you wont mistake homoplasy for homology.
Term
Why does a high mutation rate cause you to mistake homoplasy for homology?
Definition
Because when analyzing DNA if the nucleotide undergoes 2 mutations it is going to look the same as a gene that didn't mutate at all... Also long time since common descent with high rates of evolution means higher likelihod of convergent evolution.
Term
Which areas acquire mutations more rapidly: coding or noncoding regions? WHy?
Definition
Noncoding. Because natural selection has no effect on them which means mutations are neutral and can accumulate.
Term
Why are mutation rates in the 3rd position of the codon moore likely?
Definition
Because the third nucleotide in a codon usually does not change the amino acid that is produced.
Term
Why would a gene aquire mutations very slowly?
Definition
Because they perform a vital function: the histone genes are a good example.
Term
Is the rate of mutation higher in mitochondrial DNA?
Definition
Yes
Term
hat are the three reasons why rates of mutations are higher in mitochondrial DNA?
Definition
1) the DNA polymerase has no proofreeding function.
2) There are higher rates of free radicals in the mitochondria causing more mutationos.
3) there is less selective pressure on mitochondria because there are multiple mitochondria in the cell.
Term
Why are antioxidants good?
Definition
Because they take the free oxygen out of our bodies before it has a chance to affect our genomes.
Term
When would looking at the mitochondrial genome of two organisms be appropriate?
Definition
If the two organisms are thought to have evolved pretty recently.
Term
Is it better to run multiple small data sets and compare the results to choose the answer many converge on, or to run a large data set to create a tree?
Definition
Get a large data set and one tree.
Term
Why might you end up with a polytomy in a gene analysis?
Definition
Becase the species diverged so recently there had not been enough time for the relationship to resolve because there isn't enough variation.
Term
What is a species?
Definition
The smallest evolutionary unit.
Term
What is mitochondrial analysis bad at?
Definition
Determinging evolutionar relationships far back in evolutionary history
Term
Why is it easy to age hawaii?
Definition
Because the islands are the tips of volcanoes.
Term
Nuclear dna can give the evolutionary history between these time frames ...
Definition
5.1 MYA and 0.4 MYA
Term
Mt. DNA can give evolutionary history up to ... and resolve divergence from...
Definition
3.7 MYA .... 0.3-0.4 MYA
Term
T/F combining mdna and dna data sets gives the best tree
Definition
True.
Term
What has evolution showed us about the migration of drosophala flies on the hawaiian islands?
Definition
Kaua'i -> O'ahu -> 3 seperate migrations to maui -> big island. at the same time one migration from kaua'a to big island.
Term
is there an absolutely true tree?
Definition
No, they are all hypotheses.
Term
What good are phylogenetic trees?
Definition
They tell us the history of life on earth.
They help us understand the processs of evolution.
Term
can phylogenies be used to study co-evolution?
Definition
Yes.
Term
What is the biological species theory?
Definition
A species is a group of organisms that can produce viable offspring.
Term
What is the main problems with the biological species theory?
Definition
1)You must know the mating habits of the organisms in question. You can't do this with extinct organisms!
2) some species don't undergo sexual reproduction.
3) many obvious species DO hybridize and produce viable offspring. Usually these are plants.
Term
What mechanisms do species use to make sure they only mate with members of their own species?
Definition
Lock and key genetalia.
Geographical isolation
Mating rituals
different mating seasons.
Term
How do worm sperm and eggs of species like the palolo worm who put there gametes at the surface of the sea stop other species from fertilizing them?
Definition
the gametes have chemical barriers
Term
What is hybrid breakdown?
Definition
The hydbrids offspring have reduced viability or fertility.
Term
what are the three bariers to interspecies fertilization?
Definition
Prezygotic barriers
Mating barriers
Fertitization barriers
Term
What is the morphologicial species concept?
Definition
defines species based on their physical appearance.
Term
What is the problem with the morphological species concept?
Definition
only usueful for some species. protists and bacteria might look alike but are completely different.
It is often wrong
Some snails have different shells but are the same species.
Term
What is the ecological species concept?
Definition
Defines species based on where they live.
Term
What is the pleuralistic species concept?
Definition
Defines species using many different concepts: Ecological, morphological, biological...
Term
What is currently the best definition of a species?
Definition
The phylogenetic (genealogical) species concept. A species has a unique evolutionary history.
Term
what is the downside to the phylogenetic species concept?
Definition
You have to create a phylogenetic tree to define a species.
Term
when is a character said to be fixed in the population?
Definition
When ever member of the population has the exact same character.
Term
what do fixed character indicate?
Definition
reproductive isolation has occured
Term
what is population aggregation analysis?
Definition
when you examine individuals in a population to figure out if some character is fixed in the population.
It uses species identification as a criteria.
Term
what is genetic barcoding?
Definition
using a genetic sequence to identify the species.
Term
How do they do genetic barcoding?
Definition
they use the CO1 region on the mitochondrial dna
compare it to a database of CO1 sequences of barcoding studies.
Term
What is vicariance?
Definition
When a bridge prevent speciation narrows until there is geographical isolation leading to speciation
Term
what is microevolution
Definition
changes in allele frequency in a population
Term
What is macroevolution?
Definition
creation of new genera or families
Term
where does speciation sit?
Definition
On the boundary between microevolution and macroevolution
Term
What is the founder event?
Definition
When one member
Term
WHat is sympatric speciation?
Definition
When speciation occurs in a population due to different mating habits, etc
Term
what is probably the main cause of alopatric speciation?
Definition
Founder event
Term
What is alopatric speciation?
Definition
When speciation occurs due to some physical barrier
Term
What are the three stages of speciation?
Definition
1. populations have to become isolated from one another
2. traits have to diverge
3. teh divergence of traits reenforces reproductive isolation.
Term
what are the three factors that cause speciation?
Definition
Genetic drift
natural selection
sexual selection
Term
what two things can happen with allopatric speciation?
Definition
they can either remeet ancd can interbreed or they cant and speciation occcurs
Term
What is it called why hybrids are produced with reduced viability?
Definition
Reenforcement. it reenforces to barriers to gene flow.
Term
SOme hybrid lines work well together and become selected for
Definition
Yes.
Term
Hybrid zones
Definition
Areas where species overlap and hybridization is increased
Term
what habitat do viable hybrid species occupy?
Definition
the habitats between the two species they were made from
Term
What is reticulate evolution?
Definition
When hybrids have greater fitness
Term
When do the hybrid zones stabilize?
Definition
When hybrids are more fit thatn the parent species
Term
Can resluts found to be true in one specie be extrapolated to other species?
Definition
No.
Term
Who created the biological species concept?
Definition
Ernst Mayr
Term
Why is the popular misconception that "we used to be monkeys" wrong?
Definition
Because we only share a common ancestor with monkeys. Our closest living great ape relative is the pan troglodytes, the chimpanzee
Term
When and where did Homo Sapiens first originate?
Definition
100,000 years ago on the grassy plains of east Africa.
Term
Why is the misconception that man is the pinnacle of creation wrong?
Definition
Because evolution is no more slanted to give rise to humans than it is to give rise to any other animal. We are not the most recently evolved animal.
Term
What did Anaximander think man descended from in 600 BC?
Definition
A fish like creature.
Term
What is the biological junkyard hypothesis?
Definition
Animals were formed from randomly assembled arms and legs, the best combinations survived.
Term
Who thought of the biological junkyard hypothesis? When?
Definition
Empedocles (500 BC)
Term
Why is the biological junkyard hypothesis significant?
Definition
Because it was selection.
Term
Did Carol Von Linnae Believe in evolution?
Definition
No, he believed in creationism.
Term
What did Carol Von Linnae do?
Definition
Cataloged living things. Created the idea of a species.
Term
What did Bonnet believe about the origin of life?
Definition
All living things arose from germs that the planet was seeded with. These germs develop when conditions are right.
Term
What did Comte de Buffon believe about life?
Definition
There is a mold that the particles of living things were assembled in.
Term
What is spontaneous generation? What evidence supported this?
Definition
Spontaneous generation means life came into existence out of thin air. The evidence of this was that soup left for a week would be full of microorganisms. Also they thought mice spontaneously generated in bags of grain.
Term
What did Linnaeus create?
Definition
Binomial nomenclature.
Term
What information does the name Drosophila melanogaster give you?
Definition
Genus: Drosophila
Species: Melanogaster
Term
The scopes monkey trial was held when and where?
Definition
1925 in Daton TN
Term
"All scientific hypothesis must be _____" -Karl Popper
Definition
Falsifiable
Term
What is methodological naturalism?
Definition
Only natural causes can be invoked to explain natural phenomenon.
Term
What is ontological naturalism?
Definition
There are no supernatural causes, only natural causes are real.
Term
Explain intelligent design's argument using the bacterial flagella.
Definition
The bacterial flagella requires 23 separated proteins to function, the odds of evolving 23 proteins simultaneously is astronomical.
Term
In 2005 Intelligent design was deemed to be the same as ______ by the US district court in Pennsylvania.
Definition
Creationism
Term
What are the four great blows to humanity?
Definition
1. Copernicus' discovery that the earth is not the center of the universe.
2. Freud's discovery that we have an unconscious that makes us unaware of what we are doing or why we are doing sometimes.
3. The holocaust showed that man is not generally a good and kind creature.
4. Darwin showed that humans are not special, but just one of many species of animals.
Term
George Lyell (1797-1875) is called the ______ and created the theory of _______ which states ______.
Definition
Father of geology
Uniformatarianism
Geological processes taking place today are the same that took place in the distant past.
Term
Uniformitarianism leads us to the conclusion that ______
Definition
The earth is much older than previously thought.
Term
Thomas Malthus introduced the idea of overpopulation due to _____.
Definition
lack of resources.
Term
Darwin (1809-1882), a wealthy aristocrat, dropped out of ______ and couldn't find a real job. He decided to get a job on the Beagle doing what?
Definition
Medical school
He joined the Beagle in 1831, his job was to keep the captain of the Beagle company since he was also an aristocrat and there were only sailors around.
Term
What 6 factors did Darwin use to figure out evolution by natural selection
Definition
1. Old earth (Lyell)
2. Fossil record of changing organisms.
3. Species have variation.
4. Organisms seem adapted to their environment.
5. With limited resources, only some organisms will survive. (Malthus)
6. Linnaean system shows similarities between some organisms.
Term
Alfred Russel Wallace (1823-1913) did a lot of ______.
Definition
Traveling/sailing
Term
in 1852 Alfred Russel Wallace was ______.
Definition
Shipwreaked and survived 10 days on a life raft.
Term
What is the name of the famous paper written by Alfred Russel Wallace while he had malaria in 1854? Why was it significant?
Definition
On the Tendency of Varieties to Depart Indefinitely from the Original Type. It described evolution before Darwin released "On the Origin of Species."
Term
Darwin published "On the Origin of Species" in.....
Definition
1859
Term
Natural selection is competition between _______ for food.
Definition
members of the same species
Term
The _______ acts on variation in populations to produce organisms that are best adapted.
Definition
Environment
Term
Many biologists think that the first molecule to arise on earth were the ___.
Definition
Ribozymes (RNA molecules)
Term
an RNA enzyme molecule called a ribozyme was probably the first molecule because it can ____ and ____.
Definition
Catalyze certain reactions
Store genetic material
Term
Two chiral molecules are ______ of each other.
Definition
Enantiomers
Term
Two achiral molecules are ______ of each other.
Definition
diastereomers
Term
D-dopa is the enantiomer of L-dopa, does D-dopa treat parkinson's disease like L-dopa does?
Definition
No.
Term
a mixture of equal parts of two chiral molecules is called a ______
Definition
Racemic mixture.
Term
What is the evidence that the amino acids in the Murchison meteorite are not from earth or an organism?
Definition
If they were from earth there would be a racemic mixture of both enantiomers. If they were from an organism there would be just one enantiomer. There is an uneven mixture of amino acid enantiomers.
Term
Meteorites that originated on mars have been found on earth containing what kind of life?
Definition
Prokaryotes.
Term
All of the biological macromolecules that are polymers can be readily synthesized in water by ____ and broken down by _____.
Definition
dehydration synthesis
Hydrolysis
Term
Do polymer chains break down easily?
Definition
Yes
Term
How can you prevent the breakdown of polymer chains such as nucleotides?
Definition
Immobilize them on a clay-like substance. In many experiments Montmorillonite is used because many things stick to it. (it is adsorbent)
Term
Once nucleotides are adsorbed to the montmorillonite, how to you create nucleic acid polymers?
Definition
You wash the montmorillonite repeatedly with activated nucleotides
Term
How do we think protein synthesis took place in the beginning?
Definition
RNA nucleotides attached to a clay substrate bound to specific amino acids which connect together into a polymer. This polymer facilitates copying of the RNA causing the RNA to increase in number.
Term
Protobionts
Definition
Aggregations of molecules
Term
What is a liposome?
Definition
A protobiont - an aggregation of lipid molecules that forms a bubble like cell membrane.
Term
what is subduction?
Definition
When a tectonic plates slides under another tectonic plate
Term
Is graphite produced biotically or abiotically?
Definition
Abiotically and biotically.
Term
How can we tell if graphite has been produced abiotically or biotically?
Definition
the ratio of isotopes of graphite tell us whether it is abiotic graphite or biotic graphite.
Term
The oldest fossils of cells are how old?
Definition
3.4 billion years old.
Term
on the tree of life you will see many fusing branches between early organisms. This is due to _____.
Definition
Lateral gene transfer.
Term
When you look at a sedimentary rock what are the red layers? What is their significance?
Definition
Iron oxide (rust). Formed when iron and oxygen come in contact. It means there was some organism undergoing photosynthesis and pumping oxygen into the water.
Term
What happens to the oxygen produced by photosynthesis in the water?
Definition
The oxygen is taken up by the iron dissolved in the water first. This iron oxide falls to the bottom of the ocean. After all of the Iron has been used up the oxygen bubbles out of the water and enters the environment.
Term
what is oxidative phosphorylation?
Definition
the use of the electron transport chain to produce energy in eukaryotes.
Term
give examples of protists.
Definition
Paramecia and amoeba.
Term
what were probably the source of plastids?
Definition
Cyanobacteria
Term
What were probably the source of mitochondria?
Definition
Alpha proteobacteria
Term
What are ribozymes?
Definition
They are RNA molecules folded into certain shapes. They have a limited ability to do catalyze reactions.
Term
Cech and Altman found the first ribozyme in the genome of a ________.
Definition
Ciliated protist similar to the paramecium
Term
what is special about the RNA intron between the regions that code for rRNA genes in the Tetrahymena?
Definition
The RNA in these regions can splice itself out by folding itself into a tertiary structure.
Term
____ and ____ are the source of energy for almost all cellular processes
Definition
ATP
GTP
Term
Two types of RNA are used in protein synthesis _____ and ____.
Definition
tRNA and mRNA
Term
The tetrahymena ribozyme can also _____ after it has cut itself out of the genome.
Definition
Bind to other RNA molecules and cut them out by breaking the phosphodiester bond that holds adjacent nucleotides together. It then attaches this sequence to it's 3' end
Term
phosphodiester bonds _____.
Definition
hold adjacent nucleotides together
Term
What do experiments simulating conditions of an early earth show?
Definition
That Nucleotide monomers and then RNA polymers could arise outside of the cell.
Term
No experiment has shown that RNA can replicate itself, however it has been shown that RNA molecules can ______.
Definition
Replicate other RNA molecules
Term
What did the experiment using Q beta RNA and RNA replicase show?
Definition
RNA evolution. after the 74th test tube incubation *with replicase* there was a 5% difference from the original sequence.
Term
why can't proteins have been the first molecules to arise on earth?
Definition
They can't replicate themselves
Term
Why couldn't DNA have been the first molecule to arise on earth?
Definition
Because DNA needs DNA polymerase to reproduce itself.
Term
What are most enzymes made out of?
Definition
Proteins.
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