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Clemson Forest Biology Exam 1 Terms
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160
Biology
Undergraduate 2
09/13/2012

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Term
Apical Meristem 
Definition
areas of rapid cell division in shoots usually found at the tips of roots and branches; produces primary growth
Term
Lateral Meristem
Definition
areas of rapid cell growth giving rise to vascular and cork cambium; produces secondary growth
Term
Vascular Cambium
Definition
Meristematic tissue that produces xylem, phloem, and ray parenchyma 
Term
Cork Cambium
Definition
the lateral meristem that produces bark
Term
Axillary/Lateral Buds
Definition
buds formed on the sides of branches; form at the base of leaves; lateral bud growth results in production of a branch
Term
Terminal Bud
Definition
masses of meristematic tissue that may give rise to primary growth that occur on the ends of shoots and are covered and protected by cataphylls 
Term
Apical dome
Definition
the part of the apical meristem between the leaf primordia 
Term
Protoderm
Definition
Primary meristem that gives rise to cork cambium
Term
Ground meristem 
Definition
an area of primary merisematic tissue that develops into the pith and cortex and ermerges from behind the apical meristem 
Term
Procambium
Definition
Primary meristem that gives rise to vascular cambium
Term
Primary growth
Definition
Growth that increases height of tree or twig length; includes twig formation and elongation, and leaf production
Term
Secondary growth
Definition
Growth that increases the diameter of the tree through lateral meristematic activity
Term
Xylem
Definition
vascular tissue primarily responsible for carrying water and minerals; made up of vessel elements and tracheids; forms to the inside of the vascular cambium
Term
Phloem
Definition
vascular tissue that transports carbohydrates; forms to the outside of vascular cambium
Term
Cataphylls
Definition
bud scales; a type of simple leaf that surround and protect the tiny, preformed secondary leaves
Term
Epicormic sprout
Definition
shoots that grow from adventitious buds that lie dormant in stems or branches and can sprout during times of stress; aka water sprouts
Term
Terminal leader
Definition
the uppermost, upwardly growing stem of a tree; in excurrent growth, its position is maintained by hormones that supress the growth of other lateral branches
Term
Apical dominance
Definition
the phenomenon of the terminal leader maintaining upward orientation and more rapid growth than lateral branches
Term
Secondary Needles
Definition
needles borne following the growth of primary needles that usually occur in fascicles; needles produced for the remainder of the trees life are secondary
Term
Leaf primordia 
Definition
embryonic leaves that develop as a result of the apical meristem being "sloughed off" the apical dome to develop on the sides of the apical meristem
Term
Stipules
Definition
modified, rudimentary leaves found at the point of attachment of regular leaves to the stem; contribute carbon to spring growth before leaves have fully expanded
Term
Primary needles
Definition
first leaves borne after cotelydons emerge; occur singly even though needles occur in flushes later
Term
Cuticle 
Definition
waxy coating on the outside of leaves and bark that prevents desiccation
Term
Epidermis
Definition
outermost layer of cells underlying the upper and lower leaf surfaces; the outermost layer of cells underlying bark cuticle
Term
Sclerenchyma
Definition
hardened cells that provide structural support in leaves and other parts of the tree
Term
Palisade cell
Definition
columnar cells located beneath epidermis of leaves containing many chloroplasts 
Term
Spongy Mesophyll
Definition
irregular-shaped cells containing many chloroplasts; located adjacent to lower epidermis of leaf
Term
Guard cell
Definition
pairs of cells surrounding stomata that regulate the size of such structures on the surface of leaves
Term
Stomata
Definition
pore on leaf surface where most gas exchange between interior leaf tissue and its environment take place
Term
Resin duct
Definition
pockets of pitch (sap-like substance) that helps to repel bugs that feed on conifer needles
Term
Endodermis
Definition
a ring of dermal tissue surrounding the vascular bundle
Term
Transfusion tissue
Definition
a band of live and dead cells surrounding vascular tissue in pine needles
Term
Chlorenchyma
Definition
chloroplast-containing cells in some conifer needles
Term
Hypodermis
Definition
a layer of thick-walled cells below the epidermis
Term
Stem unit
Definition
stem + leaf primordia; a collective term for one leaf (in angiosperms) or one fascicle (in gymnosperms)
Term
Internode
Definition
a portion of a twig inbetween two leaves (in angiosperm) or two fascicles (in gymnosperms)
Term
Fixed growth
Definition
new stem units are formed and overwinter in buds until they experience bud break the next year in the spring; elongation occurs on a year by year basis - one flush (whorl) per year; number of leaves in a shoot are determined the year before; Determinate growth pattern - sets true terminal bud
Term
Free growth
Definition
stem units are formed while elongation is taking place; tree will keep making leaves while conditions are favorable
Term
Recurrent growth
Definition
fixed growth except terminal bud can experience bud  break in the same growing season if conditions are favorable; shoot elongation consists of more than one period of elongation and pause in primary growth in a growing season
Term
Determinate growth
Definition
true terminal bud forms near end of growing season; associated with fixed growth
Term
Indeterminate growth 
Definition
no true terminal bud forms; twig elongation and bud formation continue until stopped by frost, part of twig after last lateral bud then dies and that lateral bud becomes the terminal bud for the next growing season; associated with free growth
Term
Pseudoterminal bud
Definition
last lateral bud formed during indeterminate growth season; acts as terminal bud for the start of the next growing season
Term
Lammas growth
Definition
premature bud break/elongation of a terminal bud that usually results from a lot of rain and/or warm weather late in the growing season; usually killed by frost and causes a loss of apical dominance for that shoot
Term
Prolepsis
Definition
late season elongation of lateral buds
Term
Foxtailing
Definition
excessively strong apical dominance observed in pines planted in tropical regions resulting in long unbranched shoots and poor wood quality
Term
Stipule scar
Definition
visible wound remaining on bark of tree after stipule falls from twig
Term
Lenticle
Definition
pores for gas exchange on young twigs 
Term
Leaf scar
Definition
remnant left by fallen leaf on twig at its point of attachment 
Term
Bundle scar
Definition
visible remnant of leaf connection to vascular tissue on twig; completely enclosed by leaf scars
Term
Bud scale scar
Definition
band formed around twig as a remnant of terminal bud
Term
Phyllotaxy
Definition
the arrangement of leaves on a stem
Term
Opposite phyllotaxy
Definition
buds are arranged on twig in an orientation opposite of one another
Term
Alternate phyllotaxy
Definition
bud arrangement on twig with orientation of buds alternating from left to right or vice versa
Term
Blade
Definition
the planear portionof leaves in angiosperms
Term
Petiole 
Definition
stem-like structure attaching leaf blade to twig
Term
Midrib (rachis)
Definition
central rib of leaf that is usually continuous with petiole 
Term
Angiosperm
Definition
"hidden seed"; taxonomic class of trees in which the ovule, which after fertilization becomes the mature seed, is encased within an ovary; aka deciduous trees for losing leaves during or before the onset of winter; aka hardwood even though wood is often softer than softwood trees 
Term
Gymnosperm
Definition
"naked seed"; seeds not enclosed in an ovary and ovules exposed at time of pollination; aka evergreens because some leaves (needles) are kept through at least one winter; aka softwood because strong yet light wood good for use in construction industry
Term
Excurrent
Definition
growth form where the terminal leader grows at a faster rate than lateral branches and hormonally inhibits growth of lower branches resulting in a conical tree shape; usually occurs in conifers and some hardwoods
Term
Decurrent
Definition
growth form with a spreading crown shape resulting from multiple forking of the terminal leader and growth rates of lateral branches as fast or faster than terminal leader
Term
Meristem
Definition
area of rapid cell division and expansion
Term
Secondary xylem
Definition
xylem formed by the vascular cambium; secondary xylem forms practically all wood
Term
Tree
Definition
a woody plant that is 20+ feet at maturity with a single, unbranched trunk for several feet and a somewhat well-developed crown
Term
Monocot
Definition
subdivision of angiosperm trees in which older portion of trunk does not continue to increase in diameter as tree grows; most common are the palms
Term
Dicot
Definition
subdivision of angiosperms in which seeds have two cotyledons or seed leaves that emerge after germination; most angiosperms fall into this category
Term
Root suckers
Definition
shoots that arise from adventitious buds on roots; usually arise when injury has been done to the tree such as being felled; or can be a major form of asexual reproduction for some species
Term
Secondary Phloem
Definition
phloem produced by the vascular cambium; forms a cylinder immediately inside the bark
Term
Central zone
Definition
cells located in the top, central section of the meristem that divide infrequently and are the self-renewing undifferentiated stem cells
Term
Peripheral zone
Definition
region surrounding the central zone; cells here divide much more rapidly and become cells that contribute to the organs of the plant
Term
Chloroplast
Definition
where photosynthesis takes place; organelle of leaf cells
Term
Vein
Definition
vascular bundles
Term
Cortex
Definition
bark tissue underlying phelloderm
Term
Pith
Definition
central portion of twig or root
Term
Stem fiber
Definition
narrow and tough cells that function as support in angiosperms; xylem cells formed in addition to tracheids and vessel elements
Term
Phelloderm
Definition
living, inner bark cells that store starch
Term
Phellum
Definition
cork cells produced by the phellogen/cork cambium
Term
Ray parenchyma
Definition
radially orientated plates of living cells that are a part of secondary xylem; store and transport carbohydrates and other compounds
Term
Growth ring
Definition
a year's worth of secondary xylem growth; distinguishable because of transition from latewood to earlywood at the start of each new year
Term
Earlywood
Definition
formed in springtime when tree diameter is rapidly expanding; cells are large, of low density, and have thin cell walls 
Term
Latewood
Definition
small, high density cells with thick cell walls; forms after earlywood in the growing season
Term
Diffuse porous 
Definition
type of wood in angiosperms; small, scattered vessels formed in climates with constant water available throughout seasons
Term
Ring porous 
Definition
type of wood in angiosperms; large, concentrated vessels all formed at same time; formed for plants living in climates with seasons of more and less water
Term
Vessel elements
Definition
short, wide xylem cells adapted for transporting large amounts of water
Term
Tracheids
Definition
xylem cells that serve as both conductive and support cells in gymnosperms; in angiosperms, vessel and fiber cells are formed in addition to tracheids for xylem tissue
Term
Pit
Definition
intercellular connection where secondary cell wall is absent but primary cell wall is present
Term
Cavitation
Definition
during times of drought xylem cells can be destroyed when air bubbles form creating a vacuum because not enough water is available to move through cells
Term
Stratification
Definition
artificial method for breaking seed dormancy involving storing seed in cool, moist conditions with temperatures just above freezing to mimic chilling
Term
Scarification
Definition
artificial method for breaking seed dormancy involving mechanically or chemically breaking seed coat
Term
Chilling
Definition
certain number of hours required by some plant species at a certain temperature to break seed dormancy
Term
Mycorrhizae
Definition
symbiotic relationship between roots and fungi; fungi break down organic matter for nutrients; roots return some sugars made through photosynthesis
Term
Ectomycorrhizae
Definition
fungi located on outside of fine roots; breaks down nitrogen; more common in colder regions; macroscopic
Term
Endomycorrhizae
Definition
microscopic fungi located on inside of fine root; collects phosphorus
Term
Rhizobia
Definition
nitrogen-fixing bacteria that inhabit root nodules of legumes
Term
Actinomycetes
Definition
bacteria that form symbiotic relationships with plant; in exchange for sugars produced through photosynthesis, the bacteria provides resistant to diseases
Term
Polysaccharide
Definition
cellulose - gives structure to cell walls
Term
Primary cell wall
Definition
outermost layer of cell wall
Term
Cellulose
Definition
gives structure to cell walls
Term
Microfibril
Definition
basic structural units of cell walls; composed of cellulose
Term
Hemicellulose
Definition
less complex than cellulose but also found in plant cell walls
Term
Lignin
Definition
compound that fills the empty spaces in cell walls and binds cells together; also gives strength to cell walls
Term
Secondary cell wall
Definition
made up of three distinct layers with different orientations of cells; contains mostly cellulose
Term
Middle lamella
Definition
layer that bonds cells together; rich in lignin
Term
Plasmodesmata
Definition
channels between cell walls through which plant cells communicate and transport                                             
Term
Lumen
Definition
empty cavity insdie dead, mature fiber cells
Term
Seed cone
Definition
female, seed producing cone in conifers
Term
Microsporophyll
Definition
cone scale of male pollen cone
Term
Reaction wood
Definition
abnormal secondary xylem cells that grow in response to leaning or bending of a stem; usually results in decreased wood quality and eccentric growth rings
Term
Compression wood
Definition
reaction wood in which larger growth rings (latewood) on underside of leaning stem form to push upright; lignin rich - very strong; occurs in conifers
Term
Tension wood
Definition
reaction wood formed in hardwoods on the upperside of leaning stem to pull upright; less lignin - uses tension to pull upward; larger (latewood) rings form on top side
Term
Sapwood
Definition
wood nearest the bark that is younger and lighter in color; composed of active xylem (10% living cells), live storage cells and ray parenchyma (for storage and movement of compounds towards heartwood) 
Term
Heartwood
Definition
wood nearest the center of the trunk; oldest wood and often darker in color (more valuable); composed completely of dead secondary xylem cells and used for storage of resins and other anti-fungal substances; plays no role in conducting water or minerals 
Term
Tylose
Definition
ball of gum or resin located in heartwood to protect against fungi
Term
Ovule
Definition
rounded outgrowth of the ovary that develops into a seed after fertilization
Term
Pollen cone
Definition
male, pollen producing cone in conifers
Term
Microsporangium
Definition
Term
Microsporangium
Definition
location in male pollen cone where pollen grains are formed 
Term
Microgametophytes
Definition
pollen grains
Term
Megasporophyll
Definition
seed scale on female seed cone; covered by integument
Term
Megasporangia
Definition
two per megasporophyll; develop into megagametophytes
Term
Micropyle
Definition
small opening through which pollen tube penetrates the ovule
Term
Seed coat
Definition
outer, protective layer of a seed
Term
Integument
Definition
the outer layer of an ovule
Term
Monoecious 
Definition
separate male and female reporductive structures on the same tree
Term
Dioecious
Definition
male and female reproductive structures on different trees
Term
Complete flower
Definition
has all flower parts - corolla (petals), sepals, stamens, and pistil 
Term
Incomplete flower
Definition
missing one of the four flower parts - either corolla (petals), sepals, stamen, or pistil
Term
Perfect flower
Definition
have functioning male and female parts
Term
Imperfect flower
Definition
lack a stamen or pistil and are therefore always incomplete
Term
Pistillate 
Definition
imperfect flowers that have a pistil only
Term
Staminate 
Definition
imperfect flowers that only have a stamen
Term
Corolla
Definition
petals; inner set of floral leaves consisting of fused or separate petals that surround the carpels 
Term
Calyx
Definition
full collection of sepals on an individual flower
Term
Perianth
Definition
consists of calyx (all sepals) and corolla (all petals) on a flower; or if no sepals, perigone (all the tepals) 
Term
Stamen
Definition
the pollen producing part of a flower supported by a filament
Term
Anther
Definition
the part of the stamen where pollen is produced
Term
Filament
Definition
thin supportive stalk of the stamen 
Term
Pistil
Definition
the ovule-producing (seed-producing) structure of a flower; consists of the style, stigma, and ovary containing the ovule
Term
Stigma
Definition
the part of the pistil where pollen germinates
Term
Style
Definition
supportive structure in pistil connecting stigma to ovary
Term
Fruit
Definition
the mature ovary of a flower
Term
Endocarp
Definition
the innermost layer of the pericarp (fruit wall)
Term
Mesocarp
Definition
the middle layer of pericarp (fruit wall)
Term
Exocarp
Definition
outer layer of pericarp (fruit wall)
Term
Radicle
Definition
the root of a newly emerged seedling
Term
Cotyledons
Definition
embryonic leaves of a plant; will become the first leaves following emergece from the seed coat
Term
Hypocotyl
Definition
embryonic stem; part of seedling connecting cotyledons to radicle
Term
Epicotyl
Definition
portion of seedling above cotyledons
Term
Endosperm
Definition
seed tissue containing stored food including fats, carbohydrates, and proteins
Term
Seed viability
Definition
proportion of seed that is physiologically capable of producing a seedling
Term
Plumule
Definition
first bud of the seedling/embryonic plant; consists of the epicotyl and cotyledons
Term
Seedling sprout
Definition
regrowth of seedling if damagaed or cut back
Term
Layering
Definition
formation of roots on branches that touch the ground but are still connected to the parent tree
Term
Clone
Definition
offspring with identical genes to parent
Term
Cuttings
Definition
form of asexual plant propogation where a twig is cut from parent plant and buried in the ground except for a portion sticking out of the ground; buds sprout to form new shoots and roots develop from adventitious buds
Term
Grafting
Definition
asexual plant reproduction; fusing the scion (shoot) of one plant to the stock (rootsystem) of another to combine desirable traits of both; offspring will have no characteristics of the stock
Term
Superior tree
Definition
tree with desirable phenotypic characteristics - straight, clear bole, small crown, disease resistance, large size; cuttings taken for establishment of orchards
Term
Graft union
Definition
the point where the scion was joined with the stock; is sometimes clearly visible on older trees
Term
Micropropagation
Definition
form of asexual propogation using small tissue fragments to make a tissue and bud culture
Term
Somatic embryo
Definition
produced through somatic embryogenesis (a method of micropropogation) in an unlimited number from a single seed that will grow into identical trees; "somatic" means the embryos are created asexually
Term
Adventitious buds 
Definition
similar to dormant buds except cannot be traced to the pith; examples are root suckers or stump sprouts;  generally form on older portions of the tree; also may form when injured or when roots are exposed to light
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