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| Observable response of organisms to external or internal stimuli |
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| The study of how organisms interact with other organisms and their environment |
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| The study of processes that occur within living organisms |
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| use of specially designed signals or displays to modify behavior of others |
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| studied honey bee communication and sensory biology. the first person to demonstrate that nonhuman animals have the ability to encode and transmit spatial information. "the waggle dance" |
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| Tactile, Chemical, Auditory, Visual |
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| often used to convey information about food resources |
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| often critical in species identification during mating |
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| often critical in species identification during mating |
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chemicals to which special receptor cells may be attuned >vomeronasal or jacobson's organ |
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| Vomeronasal / jacobson's organ |
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| special organ used to detect pheromones |
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| any modification of behavior that results from experience rather than maturation |
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| learning in which a behavior is changed or conditioned through association |
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| Classical conditioning (pavlovian) |
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| An involuntary response becomes associated with an unrelated stimulus |
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| Operant conditioning (skinnerian) |
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| behavior is reinforced by positive and negative consequences |
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| learning that is not associated with a particular stimulus (ie Habituation) |
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learning in which there is a decrease in the response to a stimulus due to repetition form of nonassociative learning |
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| solving problems with thought rather than environmental feedback |
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| instinct or innate behavior |
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heritable and genetically specified behavior performed perfectly the first time |
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sign stimulus or releaser
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| any simple signal that elicits a specific behavioral response, known as a fixed action pattern |
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| fixed action pattern (FAP) |
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| a specific behavioral in response to a sign stimulus |
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hygenic strain vs non hygenic strain found that one gene controlled for uncapping a cell, one gene for controlled larvae removal showed the effect of a single gene on complex behavior |
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| examples of interaction between genetics and learning |
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birds songs if bird doesn't hear song during critical period, it will song an abnormal song. if bird hears song of different bird species, will sing abnormal song but if hears song sung by bird of own species at critical period, will sing correct song |
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| proximate causation of animal behavior |
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how does the behavior work? we look at: internal physiology (processes that occur within living organisms) different than the ecology or ultmate causation (why?) |
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| ultimate causation of animal behavior |
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why has behavior evolved? we look at the adapative value, or how the animal's survival or reproductive success has been influences different than the proximate causation- how does the behavior work |
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| The ways in which behavior serves as an adaptation and allows an animal to increase its survival and reproductive success |
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| natural selection results in foraging behavior that is as energetically efficient as possible |
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| When might optimal foraging theory not predict foraging behavior? |
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| Periods of special nutrient needs, periods of heavy predation (or other tasks) |
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| The maintenance of an area used by an individual (to the exclusion of other individuals) |
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| portion of home range defendable against comspecifics and other species |
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| What does territory provide? |
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| food, protection, reproductive needs (mates, breeding sites) |
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| complete and sufficient territory- evenly distributed food resources |
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| Used for mating and nesting but not for feeding- sparse patches of food |
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| Mating territory only- leks, unlimited food |
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| an area divided into a cluster of territories held purely for mating, nothing else provided. sage grouse |
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| feeding territory only- limited food, easily defended |
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| limits on female fecundity |
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| limited by resources, so therefore, females should be very selective, provide for offspring, and their distribution should be dictated by resources |
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| limited by number of females, thus distribution dictated by female distribution |
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| microevolutionary process that favors traits that give individual a competitive edge in attracting and holding mates. results in evolution of secondary sex characteristics. |
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| one male mates with one female |
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| one male mates with more than one female |
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| one female mates with more than one male |
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| monogamy, polygyny, polyandry, promiscuity |
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| how synchrony of female receptivity influences males access to mates: |
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if females become receptive at different times of the year, males can mate sequentially with multiple females (polygyny) if females are receptive at only one time of the year, males have little time for multiple mates (monogamy) |
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| how mating systems depend on offspring life history patterns: |
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if altricial, monogamous if precocial, polygynous |
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| long-range movement of a population associated with a change in season |
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| how do migrating birds find their way? |
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navigation- mature birds can correct their course when moved and find traditional grounds orientation- young birds continue to fly in same direction after being moved without adjusting for the movement |
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| a group of organisms of the same species that are organized in a cooperative manner |
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"truly social" 1. division of labor in reproduction 2. cooperative care of brood 3. overlap of generations |
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| the performance of an action that benefits another individual at the cost of the individual performing the behavior |
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natural selection that favors "altruism" directed towards close relatives "i would willingly lay down my life for two of my brothers or eight of my first cousins" |
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| coefficients of relatedness |
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siblings, parents r=.5 grandparents, grandchildren, uncles r=.25 cousins=.125 |
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an allele that contributes to altruistic behavior can increase in frequency in a population if Br>C C=cost B=fitness benefit r=coefficient of relatedness |
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enhances effects of kin selection which increases probability of eusociality females are diploid, males are haploid so offspring are .75=r related to mothers |
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| groups of interbreeding individuals occupying the same habitat at the same time |
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| the study of how populations grow and what factors promote and limit growth |
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| clumped pattern of spacing |
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most common, uneven resources benefits: (plants: wind break, favorable microclimate) (animals: locating food, surviving predator attacks) |
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| random pattern of spacing |
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| uncommon, uniform resources, low competition |
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| uniform pattern of spacing |
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| uncommon, uniform resources, high competition |
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the measurement of a population helps to predict how populations will change in future |
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| group of individuals of the same age |
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| percentage of an original populatin that survives to a given age |
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| set of interacting traits that allocate resources to different components of fitness: fecundity, survival and timing of events crucial to fitness |
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| how unstable environments affect life history strategy |
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| mortality is unavoidable, so doesn't do much good to delay reproduction because getting large doesn't matter in the end |
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| how stable environment affects life history strategy |
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| mortality is avoidable, so there's greater selective pressure to get large and delay reproduction |
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| dioecious sexual reproduction |
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male and female structures on different individuals "two house" |
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| monoecious sexual reproduction |
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hermaphroditic male and female sexual structures on the same individual "one house" |
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alternate hermaphrodites sex changes with environemental conditions jack-in-the-pulpit |
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| benefits and costs of hermaphroditism |
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benefits: increases mating possibilities, decreases inbreeding costs: really expensive to be both male and female |
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| development of egg without fertilization |
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| environmental factors for parthenogenesis |
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high latitudes, high altitudes, rapid temperature change, freshwater (vs marine) why? more difficult to find mates, need to reproduce rapidly when conditions are good |
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| costs: 50% of genes aren't passed on, slower population growth, may be dependan on pollinators, sexual selection equipment very expensive, mate selection and guarding, stds |
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| sexual organisms can adapt to the demands of their environments because they produce a variety of progeny, recessive genes are masked, and it feels so good |
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an organism that lives in or on another organism (host) from which it aquires nutrients most parasites are host-specific. low lethality, high intimacy |
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all resources focused on reproduction in one short span of time, then dies found in short lives species in which the cost of staying alive between broods is great, when fecundity entails large survivorship costs r-selected species? |
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produce offspring several times over many reproductive seasons k-selected species? altricial? |
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| number of offspring produced by a particular reproductive event |
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| Predator Satiation Hypothesis |
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synchrony of of emergence must be very precise in order to flood predatos at the same time emergence must not occur frequently, to prevent predators from cueing in |
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| what factors determine population growth? |
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| fecundity, mortality, immigration, emigration |
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| as r increases, a population grows more rapidly |
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| exponential growth formula |
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ΔN =(b-d)N Δt N=number of individuals in population t=time b=average birth rate per individual in population d=average death rate per individual in population if b>d population grows, b<d declines, b=d stays same |
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maximum density of individuals that the environment can support |
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| density-dependent limitation |
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carrying capacity of the environment limits population growth become more severe as k is approached |
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| density-independent growth |
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| growth continues indefinitely because increases in the size of the population do not affect the per capita rate of increase |
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| number of species in a community |
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| a measure of the number of species and their relative abundance |
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| simple v. complex communities |
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simple communities tend to have less diversity, fewer species complex communitites more diversity and more species |
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| diversity-stability hypothesis |
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| species rich communities are more stable than species poor communities |
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| intermediate disturbance hypothesis |
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| moderately disturbed communities are more diverse |
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| control community composition and maintains diversity |
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| orderly process of temporal change in a community's composition |
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| alterations initiated by pioneering communities on newly formed land |
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| establishment of a community after the original community was destroyed |
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a model used to explain succession colonizing species make the environment a little more suitable for other species |
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a model used to explain succession colonizing species make the environment less suitable for others |
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a model used to explain succession new colonists are neither inhibited nor facilitated by previous colonists |
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| primary succession along the lake michigan shoreline |
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| from coast to inland, shows stages of succession in order of disturbance levels |
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| the oldest and most stable stage of succession |
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| the effect of two species or two individuals trying to use the same limited resource |
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| intraspecific competition |
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| members of same species competing for resources. major density-dependent limitation |
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| interspecific competition |
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| the extent of competition between species is dependent on the density of the competing species relative to the shared resources |
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| gause's principles of competition |
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| species can coexist if they don't occupy same niche |
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the ways in which an organism adapts to environment to acquire essential resources |
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| the niche an organism CAN occupy |
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| the niche an organism does occupy- not neccessarily the fundamental niche |
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| rapid divergent evolution in sympatric species of characters that can help minimize competition |
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| two or more species living together |
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one species benefits, the other recieves no benefits and no harm sea anemone and clownfish |
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| one species benefits and individuals of another species are harmed |
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variation is needed to compete with a rapidly evolving background of ever changing environmental conditions: predators, parasites, prey "now, here you see, it takes all the running you can do to keep in the same place" |
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| insects that lay their eggs on living hosts which are eventually killed |
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